Daxishan, Linglongta (Jurassic to of China)

Also known as Wukongopterus, Darwinopterus type locality, Linglengta

Where: Liaoning, China (40.9° N, 120.0° E: paleocoordinates 42.1° N, 123.2° E)

• coordinate stated in text

• local area-level geographic resolution

When: Tiaojishan Formation, Late/Upper Bathonian to Late/Upper Bathonian (167.7 - 150.8 Ma)

• Wang et al. (2009) referred to this locality as from the Daohugou Formation (?Late Jurassic-Early Cretaceous), while Lü et al. (2009) suggested that it is the Tiaojishan Formation (Middle Jurassic).

•"The Tiaojishan Formation is generally regarded as Middle (Zhang & Zheng 1991; Liu et al. 2006; Cheng & Li 2007) to Upper Jurassic (Chang et al. 2009) (Late Bathonian-Early Kimmeridgian) in age, although it may date to the early Middle Jurassic (Aalenian) in Jianchang County (Davis et al. 2001). The unit has been dated using radiometric techniques at 169-152 Mya by Liu et al. (2006) and 165-156 Mya by Zhang et al. (2008)." (Lü et al. 2010: supplement)

• bed-level stratigraphic resolution

Environment/lithology: lacustrine; tuffaceous, gray sandstone and gray, green mudstone

• "The muddy shales and mudstones, and the tuffaceous layers, have yielded an important fossil assemblage" (Lü et al. 2011: supplement) Bed 13: "Grayish-green and shallow grayish mudstone and muddy shale, occasionally tuffs and tuffaceous sandstone…fossils…were unearthed from tuffaceous sediments or tuff in the lower parts of this bed."

Size class: macrofossils

Preservation: adpression, soft parts

Reposited in the IVPP

Collection methods: quarrying, surface (in situ), mechanical,

• ZMNH: Zhejiang Museum of Natural History. YH: Yizhou Museum, Yixian, Liaoning.

•PMOL: Paleontological Museum of Liaoning, Shenyang Normal University.

•Fossils collected primarily by local farmers

•The heterodontosaurid ornithischian Tianyulong was originally reported to be from the Yixian Formation (Early Cretaceous) of Jianchang County, but is now believed to be from Linglongta (Lü et al. 2011: supplementary data)

Primary reference: X. Wang, A. W. A. Kellner, S. Jiang and X. Meng. 2009. An unusual long-tailed pterosaur with elongated neck from western Liaoning of China. Anais da Academia Brasileira de Ciências 81(4):793-812 [R. Butler/R. Butler]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 92421: authorized by Richard Butler, entered by Richard Butler on 11.11.2009, edited by John Alroy, Matthew Carrano, Jonathan Tennant and Emma Dunne

Creative Commons license: CC BY (attribution)

Taxonomic list

• It is now known where in this area the Xiaotingia holotype was found, and it could have come from the Yixian Fm., which also outcrops in the area.
Insecta
  -
Insecta indet.10 Linnaeus 1758 insect
Branchiopoda
 Diplostraca - Euestheriidae
Euestheria sp.10 Depéret and Mazeran 1912 clam shrimp
Ostracoda
 Podocopida - Limnocytheridae
 Podocopida - Darwinulidae
Bivalvia
 Cardiida - Shaanxiconchidae
Mammalia
 Theriamorpha -
Juramaia sp.11 Luo et al. 2011 eutherian
Reptilia
 Testudines -
Testudines indet.3 Batsch 1788 turtle
PKUP V1070
 Ornithischia -
Tianyulong confuciusi n. gen. n. sp.16
Tianyulong confuciusi n. gen. n. sp.16 Zheng et al. 2009 ornithischian
STMN 26-3, incomplete skull and postcranium with filamentous integumentary structures
 Theropoda - Troodontidae
Xiaotingia zhengi n. gen. n. sp.14, Anchiornis huxleyi n. gen. n. sp.15
Xiaotingia zhengi n. gen. n. sp.14 Xu et al. 2011 maniraptoran
Anchiornis huxleyi n. gen. n. sp.15 Xu et al. 2009 maniraptoran
IVPP V14378, BMNHC PH828
 Pterosauria - Istiodactylidae
Archaeoistiodactylus linglongtaensis n. gen. n. sp.5 Lü and Fucha 2010 pterosaur
JPM04-0008, partial skeleton
 Pterosauria - Wukongopteridae
Kunpengopterus antipollicatus n. sp.17 Zhou et al. 2021 pterosaur
Holotype: BPMC 0042.
Kunpengopterus sinensis n. gen. n. sp.13 Wang et al. 2010 pterosaur
IVPP V16047 (holotype), nearly complete skeleton
 Pterosauria - Scaphognathidae
Jianchangnathus robustus n. gen. n. sp.2 Cheng et al. 2012 pterosaur
IVPP V16866, partial skeleton with complete skull and lower jaw
cf. Jianchangnathus robustus3 Cheng et al. 2012 pterosaur
BMNHC PH000988
Fenghuangopterus lii n. gen. n. sp.6 Lü et al. 2010 pterosaur
CYGB-0037 (holotype), incomplete skeleton with a partial skull and lower jaws
 Pterosauria - Rhamphorhynchidae
Scaphognathus robustus n. sp.1 Cheng 2012 pterosaur
Holotype: IVPP V16866, incomplete articulated skeleton. Holotype: IVPP V16866, incomplete articulated skeleton.
Jianchangopterus zhaoianus n. gen. n. sp.4 Lü and Bo 2011 pterosaur
YHK-0931, complete skeleton
 Pterosauria -
Changchengopterus pani18 Lü 2009 pterosaur
PMOL-AP00010, articulated postcranial skeleton
Darwinopterus sp.7 Lü et al. 2010 pterosaur
ZMNH M8802, complete skeleton with egg
Darwinopterus modularis n. gen. n. sp.8 Lü et al. 2010 pterosaur
ZMNH M8782 (holotype), skull and partial postcranium of a putative adult. YH-2000, almost complete skeleton of a putative female
Darwinopterus robustodens n. sp.9 Lü et al. 2011 pterosaur
41HIII-0309A
Darwinopterus linglongtaensis n. sp.13 Wang et al. 2010 pterosaur
IVPP V16049 (holotype), complete articulated specimen
Wukongopterus lii n. gen. n. sp. Wang et al. 2009 pterosaur
IVPP V15113 (holotype), nearly complete skeleton
Douzhanopterus zhengi n. gen. n. sp.12 Wang et al. 2017 pterosaur
STM 19–35A and B
Amphibia
 Caudata - Urodela
Urodela indet.10 Duméril 1805 salamander
Chunerpeton tianyiensis11 Gao and Shubin 2003 salamander
Osteichthyes
 Ptycholepiformes - Ptycholepididae
"? Ptycholepidae indet." = Ptycholepididae10
"? Ptycholepidae indet." = Ptycholepididae10 Brough 1939 ray-finned fish
 Bennettitales -
Zamites cf. gigas10 Lindley and Hutton 1831
Polypodiopsida
 Equisetales - Equisetidae
 Cyatheales - Polypodiidae
Coniopteris margaretae10 scaly tree fern
Coniopteris hymenophylloides10 Brongniart 1828 scaly tree fern
Ginkgoopsida
 Ginkgoales -
 Czekanowskiales -