Paali Nala level C2 (DBC2) (Oligocene of Pakistan)

Also known as Bugti Hills; Gandoï syncline; DB-C2; Dera Bugti C2

Where: Balochistan, Pakistan (28.9° N, 69.1° E: paleocoordinates 19.0° N, 67.8° E)

• coordinate stated in text

• small collection-level geographic resolution

When: Bugti Member (Chitarwata Formation), Early/Lower Oligocene (33.9 - 28.4 Ma)

• "in the lowermost levels of the Lower Chitarwata Formation, which is early Oligocene in age"

• bed-level stratigraphic resolution

Environment/lithology: fluvial-deltaic; sandstone

Size classes: macrofossils, mesofossils

Collected in 2000 - 2004

Collection methods: bulk, sieve,

• material is in the collection of the Institut des Sciences d l'Evolution, Université de Montpellier II; collections were made by "intensive screen-washing"

Primary reference: L. Marivaux, P.-O. Antoine, S. R. H. Baqri, M. Benammi, and Y. Chaimanee. 2005. Anthropoid primates from the Oligocene of Pakistan (Bugti Hills): Data on early anthropoid evolution and biogeography. Proceedings of the National Academy of Sciences 102(24):8436-8441 [J. Alroy/J. Alroy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 78520: authorized by John Alroy, entered by John Alroy on 07.02.2008, edited by K. Christopher Beard, Mark Uhen, Pauline Coster and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

Mammalia
 Perissodactyla - Rhinocerotidae
Epiaceratherium cf. magnum2 Uhlig 1999 rhinoceros
DBC2-RHI-1-3
 Artiodactyla - Suidae
Hyotherium sp.11 von Meyer 1834 pig
UMC-DBC 2242 and 2244
 Artiodactyla - Tayassuidae
Pecarichoerus sp.11 Colbert 1933 peccary
UMC-DBC 2240 and 2241
 Artiodactyla -
Ruminantia indet.9 ruminant
DBC2 5037
Mosaicomeryx aff. quercyi9 Jehenne 1987 ruminant
DBC2 5040, 5043, 5158, 5160, 5172, 5173 and 5182
Paalitherium gurki n. gen. n. sp.9 Métais et al. 2017 ruminant
DBC2 5186 - holotype; referred material: DBC2 5036, 5042 and 5181
Sanitherium sp.11 von Meyer 1866 ruminant
UMC-DBC 2239
 Artiodactyla - Lophiomerycidae
Nalameryx savagei10 Nanda and Sahni 1990 ruminant
DBC2 specimens
Nalameryx sulaimani n. sp.10 Métais et al. 2009 ruminant
DBC2 5108 - type
Lophiomeryx kargilensis10 Nanda and Sahni 1990 ruminant
DBC2 5009-5011, 5028, 5029, 5034, 5035, 5037, 5193-5200
 Artiodactyla - Palaeomerycidae
aff. Amphitragulus sp.9 Pomel 1847 ruminant
DBC2 5038, 5041, 5157, 5161-5166, 5168 and 5185
 Rodentia - Aplodontidae
Oligopetes sp.6 Heissig 1979 mountain beaver
 Rodentia - Cricetidae
Atavocricetodon paaliense n. sp.4 Marivaux et al. 1999 mouse
Pseudocricetodon nawabi n. sp.4 Marivaux et al. 1999 mouse
 Rodentia - Dipodidae
Allosminthus sp.6 Wang 1985 rodent
 Rodentia -
 Rodentia - Chapattimyidae
Baluchimys chaudryi6 Flynn and Cheema 1994 rodent
Zindapiria quadricollis6 Flynn and Cheema 1994 rodent
 Rodentia -
Bugtimys zafarullahi n. gen. n. sp.5
Bugtimys zafarullahi n. gen. n. sp.5 Marivaux et al. 2002 rodent
DBC 1261 - holotype
 Rodentia - Diatomyidae
cf. Fallomus sp.6 Flynn et al. 1986 rodent
DBC 295, 296, 476-494
Fallomus ginsburgi n. sp.6 Marivaux and Welcomme 2003 rodent
DBC 683 - type
Fallomus quraishyi n. sp.6 Marivaux and Welcomme 2003 rodent
DBC 438 - type
Fallomus razae6 Flynn et al. 1986 rodent
DBC 474, 475, 1001-1259, 1277
 Rodentia -
Hodsahibia gracilis n. sp.6 Marivaux and Welcomme 2003 rodent
DBC 302 - type
Hodsahibia beamshaiensis n. sp.6 Marivaux and Welcomme 2003 rodent
DBC 1438 - type
Hodsahibia azrae6 Flynn et al. 1986 rodent
DBC 308-373
 Primates - Amphipithecidae
Bugtipithecus inexpectans n. gen. n. sp. Marivaux et al. 2005 monkey
UMC-DBC 2174 - holotype; referred material: UMC-DBC 2172, 2173, 2175-2178 and 2191
 Primates - Eosimiidae
Phileosimias brahuiorum n. sp. Marivaux et al. 2005 monkey
UMC-DBC 2221 - type; referred material: UMC-DBC 2220 and 2222
Phileosimias kamali n. gen. n. sp. Marivaux et al. 2005 monkey
UMC-DBC 2199 - holotype; referred material: UMC-DBC 2192-2219
 Primates - Sivaladapidae
Guangxilemur singsilai n. sp.8 Marivaux et al. 2002 primate
 Primates - Ekgmowechashalidae
Bugtilemur mathesoni n. gen. n. sp.7
Bugtilemur mathesoni n. gen. n. sp.7 Marivaux et al. 2001 primate
 Theriamorpha - Herpetotheriidae
Asiadidelphis akbarbugtii n. sp.3 Crochet et al. 2007 marsupial
PMNH−DBC2−2180 - type; referred material: PMNH−DBC2−2179 and 2223
Chondrichthyes
 Carcharhiniformes - Sphyrnidae
Sphyrna sp.1 Rafinesque 1810 hammerhead shark
 Carcharhiniformes - Carcharhinidae
Rhizoprionodon sp.1 Whitley 1929 sharpnose shark
Carcharhinus sp.1 de Blainville 1816 requiem shark
species 2
Carcharhinus perseus n. sp.1 Adnet et al. 2007 requiem shark
 Rhinopristiformes - Pristidae
? Pristis cf. lathami1 Galeotti 1837 sawfish
 Rhinopristiformes - Rhynchobatidae
Rhynchobatus cf. pristinus1 Probst 1877 wedgefish
 Rajiformes - Rhinobatidae
Rhinobatos sp.1 Link 1790 guitarfish
 Myliobatiformes - Dasyatidae
Dasyatis sp.1 Rafinesque 1810 whiptail stingray
 Myliobatiformes - Myliobatidae
Myliobatis sp.1 Cuvier 1816 eagle ray
 Myliobatiformes - Aetobatidae
Aetobatus sp.1 Blainville 1816 stingray
 Myliobatiformes - Gymnuridae
Himantura sp.1 Müller and Henle 1837 stingray