Khovboor (PIN coll. 3101) [SMPE] (Cretaceous to of Mongolia)

Also known as Guchin Us Somon; Khoobur; Khobur; Khoboor; Hobur; Höövör; Khoboor, Guchinus, Khovur

Where: Övörkhangai Aymag, Mongolia (42.5° N, 102.4° E: paleocoordinates 43.0° N, 100.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Dzunbain Formation, Late/Upper Aptian to Late/Upper Aptian (122.5 - 109.0 Ma)

• "Lower Cretaceous (possibly Aptian)" and from the Khukhtekskaya Svita = Dzunbayn = Dzunbain Formation according to Dashzeveg 1975, but "? Aptian or Albian" according to Wible et al. 2001. Shown in column as "Khobur" beds (Shuvalov, 1974).

•Alifanov 2019: assumed to belong to the Hühteeg Horizon without formal assignment to any formation identified in Mongolia and are dated either to the Aptian age including the Albian age (Shuvalov, 2000) or to the late Aptian age and the entire Albian age (Lucas, 2006). According to our preliminary data, the Hühteeg Horizon corresponds to the time interval that includes the late Aptian and early Albian ages.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine; poorly lithified, gray, yellow sandstone and red, blue claystone

• from illustrated section column in Shuvalov 1974: "unconsolidated and weakly cemented sandstones"

•from K & K 1974: "an alternation, sometimes rhythmic, of gray sandstones, yellowish sands, blue and reddish clays. The bones were found in interlayers of sandstones, sands, and silts."

Size classes: macrofossils, mesofossils

Collected by P. Kozlov, P. Khosbayar, SMPE, SMGE, MAE in 1926; 1964; 1969–1972, 1991, 1997, 1999; reposited in the PIN

Collection methods: bulk, sieve,

• original collections consisted of "About seven tons of sediments... have been washed and screened" and the collection "is housed in the Palaeontological Institute of the USSR Academy of Sciences" under the charge of B. A. Trofimov

Primary reference: V. F. Shuvalov. 1974. O geologicheskom stroenii i vozraste mestonakhozhdenii Khobur i Khuren-Dukh [On the geology and age of the Khobur and Khuren-Dukh localities]. In N. N. Kramarenko, B. Luvsandansan, Yu. I. Voronin, R. Barsbold, A. K. Rozhdestve (eds.), Mesozoic and Cenozoic Faunas and Biostratigraphy of Mongolia; The Joint Soviet-Mongolian Paleontological Expedition, Transactions 1:296-304 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 77191: authorized by John Alroy, entered by John Alroy on 28.12.2007, edited by Roger Benson, Matthew Carrano, Philip Mannion and Grace Varnham

Creative Commons license: CC BY (attribution)

Taxonomic list

• Dashzeveg reports that four taxa have been "named but not described" by Trofimov in Beliayeva et al. 1974: Gobioconodon borissiaki, Gobiodon infinitus, Prokennalestes kozlovi, and Prozalambdalestes simpsoni numerous later papers by Trofimov, Kielan-Jaworowska, Dashzeveg, and others report the remaining fauna.

•Kalandadze & Kurzanov 1974 list "five new forms" of Lacertilia; these are certainly among those named later and not included in the list.

unclassified
  -
Tracheophyta indet.11
"fossilized wood"
Branchiopoda
 Phyllopoda -
Phyllopoda indet.11 branchiopod
Gnathostomata
  -
Osteichthyes indet.11 bony fish
fish teeth
Amphibia
 Salientia - Discoglossidae
Eodiscoglossus sp.15 Villalta 1957 painted frog
fragments of L maxilla
Mammalia
 Multituberculata - Plagiaulacidae
Plagiaulacidae indet. Gill 1872 multituberculate
"gen. et sp. nov."
 Multituberculata - Eobaataridae
Eobaatar magnus n. gen. n. sp.13, Eobaatar minor n. sp.13, Monobaatar mimicus n. gen. n. sp.13
Eobaatar magnus n. gen. n. sp.13 Kielan-Jaworowska et al. 1987 multituberculate
PIN 3101/57 (holotype left P4) and other specimens
Eobaatar minor n. sp.13 Kielan-Jaworowska et al. 1987 multituberculate
PIN 3101/70 (holotype right mandible) and GI PST 10-23 (anterior part of a left P4)
Monobaatar mimicus n. gen. n. sp.13 Kielan-Jaworowska et al. 1987 multituberculate
PIN 3101/65 (holotype partial left maxilla), 3101/74 (left maxilla), GI PST 10-26 (?P4), 3101/61 (left M2))
 Multituberculata - Arginbaataridae
Arginbaatar dimitrievae n. gen. n. sp.13
Arginbaatar dimitrievae n. gen. n. sp.13 Trofimov 1980 multituberculate
many specimens
 Theriamorpha -
Kielantherium gobiensis n. gen. n. sp.7, Prozalambdalestes simpsoni n. gen. n. sp.11, Hovurlestes noyon n. gen. n. sp.14, Prokennalestes trofimovi n. gen. n. sp.12, Prokennalestes kozlovi n. gen. n. sp.11, Prokennalestes minor n. sp.12, "Arguimus khosbajari n. gen. n. sp." = Arguimus khosbajari8, "Arguitherium cromptoni n. gen. n. sp." = Arguimus khosbajari9
Kielantherium gobiensis n. gen. n. sp.7 Dashzeveg 1975 mammal
Including holotype, a mandible reported by Dashvezeg & Kielan-Jaworowska (1984) (GI PST 10-16), and specimens reported by Lopatin & Averianov (2007): PIN 3101/110 (upper molar), PIN 3101/32 (dentary fragment)
Prozalambdalestes simpsoni n. gen. n. sp.11 Trofimov 1974 eutherian
Hovurlestes noyon n. gen. n. sp.14 Lopatin and Averianov 2018 eutherian
PIN, no. 3101/111 - holotype
Prokennalestes trofimovi n. gen. n. sp.12 Kielan-Jaworowska and Dashzeveg 1989 eutherian
19 specimens, plus 2 specimen reported by Sigogneau-Russell et al (1992)
Prokennalestes kozlovi n. gen. n. sp.11 Trofimov 1974 eutherian
Prokennalestes minor n. sp.12 Kielan-Jaworowska and Dashzeveg 1989 eutherian
19 specimens, plus 2 specimen reported by Sigogneau-Russell et al (1992)
"Arguimus khosbajari n. gen. n. sp." = Arguimus khosbajari8 Dashzeveg 1979 mammal
GISPS 10-15 (holotype dentary fragment)
"Arguitherium cromptoni n. gen. n. sp." = Arguimus khosbajari9 Dashzeveg 1979 mammal
PS 10-31 (holotype partial dentary) reported by Dashvezeg & Kielan-Jaworowska (1998)
  -
"Gobiodon infinitus n. gen. n. sp." = Gobiotheriodon infinitus11
"Gobiodon infinitus n. gen. n. sp." = Gobiotheriodon infinitus11 Trofimov 1980 mammal
"gen. et sp. nov."
 Eutriconodonta - Gobiconodontidae
"Guchinodon hoburensis n. gen. n. sp." = Gobiconodon hoburensis10, Gobiconodon borissiaki n. sp.11
"Guchinodon hoburensis n. gen. n. sp." = Gobiconodon hoburensis10 Trofimov 1978 mammal
plus PSS 10-37a (partial right maxilla), 10-37b (partial left maxilla), 10-37c (partial right dentary), 10-21 (crown of right m5)
Gobiconodon borissiaki n. sp.11 Trofimov 1978 mammal
"gen. et sp. nov.", PSS-MAE-104, right petrosal
Reptilia
 Testudines - Lindholmemydidae
Mongolemys sp.11 Khosatzky and Mlynarski 1971 turtle
 Testudines - Macrobaenidae
Hangaiemys hoburensis11 Sukhanov and Narmandakh 1974 turtle
 Squamata -
"Iguania" indet.2 squamates
"numerous unpaired and rugose iguanian-like frontals with a clear incision on their parietal border for the parietal opening"
 Squamata - Xantusiidae
"Xantusidae indet." = Xantusiidae1
"Xantusidae indet." = Xantusiidae1 Baird 1858 night lizard
 Squamata - Paramacellodidae
Paramacellodidae indet.1 Estes 1983 squamates
Naimanosaurus dinosauroaequalis n. gen. n. sp.3 Alifanov 2019 squamates
PIN, no. 3334/516, right dentary with teeth
 Squamata -
Hoburogekko suchanovi n. gen. n. sp.6
Hoburogekko suchanovi n. gen. n. sp.6 Alifanov 1989 squamates
in Alifanov (1989); PIN 3334-500 - holotype (most of the muzzle); referred specimen: PIN 3334-503 (partial jaw)
 Squamata - Ardeosauridae
Gurvelus khangaicus n. gen. n. sp.3
Gurvelus khangaicus n. gen. n. sp.3 Alifanov 2019 squamates
Holotype: PIN, no. 3334/518 (partial skull)
 Squamata -
"Hodzhaculia sp." = Hodzhakulia1
"Hodzhaculia sp." = Hodzhakulia1 Nessov 1985 worm lizard
 Squamata - Priscagamidae
Priscagamidae indet.1 Borsuk-Bialynicka and Moody 1984 squamates
 Squamata - Dorsetisauridae
Dorsetisaurus sp.1 Hoffstetter 1967 squamates
"numerous isolated maxillae and dentaries" (Alifanov 2000)
Paradorsetisaurus postumus n. gen. n. sp.4 Alifanov 2019 squamates
Holotype: PIN 3334/517, right maxilla
 Squamata - Xenosauridae
Xenosauridae indet.1 Cope 1866 squamates
Xenostius futilus n. gen. n. sp.4 Alifanov 2019 squamates
Holotype: PIN 3334/522, left dentary PIN 3334/521; left dentary with teeth.
 Squamata - Globauridae
Khereidia aptiana n. gen. n. sp.3
Khereidia aptiana n. gen. n. sp.3 Alifanov 2019 squamates
Holotype: PIN, no. 3334/523, right dentary
 Squamata - Slavoiidae
Slavoia sp.1 Sulimski 1984 squamates
 Squamata - Eoxantidae
Palaeoxanta conicodentata n. gen. n. sp.3
Palaeoxanta conicodentata n. gen. n. sp.3 Alifanov 2019 squamates
Holotype: PIN, no. 3334/520, left dental bone; PIN, no. 3334/555, right dental bone with teeth
 Ornithischia - Ankylosauridae
Shamosaurus scutatus16 Tumanova 1983 ankylosaurid
PIN N3101
 Ornithischia -
Psittacosaurus mongoliensis Osborn 1923 ceratopsian
 Theropoda -
Theropoda indet.11 theropod
large form
 Theropoda - Dromaeosauridae
Dromaeosauridae indet.5 Colbert and Russell 1969 maniraptoran
 Saurischia -
Sauropoda indet.11 Marsh 1878 sauropod