Bore Kevelaer, depth 85m (Miocene to of Germany)

Where: North Rhein-Westfalia, Germany (51.6° N, 6.3° E: paleocoordinates 51.8° N, 4.6° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Early/Lower Miocene to Early/Lower Miocene (23.0 - 11.6 Ma)

• boundary Reinbek-Tichelh./Hemmoor stage

• bed-level stratigraphic resolution

Environment/lithology: marine; unlithified, fine-grained, brown sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1994

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 62884: authorized by Martin Aberhan, entered by Sabine Nürnberg on 01.08.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Gymnolaemata
 Cheilostomata - Membraniporidae
Reussirella haidingeri1, Discoporella umbellata1
Discoporella umbellata1 Defrance 1823
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
Polychaeta
  -
Polychaeta indet.1
Polychaeta indet.1 Grube 1850
Sedentarida indet.
 Sabellida - Serpulidae
Filograna sp.1
Filograna sp.1 Oken 1815
Scaphopoda
 Dentaliida - Dentaliidae
Antalis dumasi1, Antalis aff. exlamarcki1, "Dentalium bouei" = Antalis bouei1, Lentigodentalium pseudomutabile1, "Pseudantalis aturensis" = Dentalium (Pseudantilis)1
Antalis dumasi1 tusk shell
Antalis dumasi subsp.
"Dentalium bouei" = Antalis bouei1 Deshayes 1825 tusk shell
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
 Gadilida - Gadilidae
Cadulus aff. subfusiformis1, Gadila dingdenensis1
Gadila dingdenensis1 tusk shell
Gastropoda
 Entomotaeniata - Cimidae
Cima neglecta3, Actaeopyramis elatus3
 Heterostropha - Mathildidae
"Mathilda (Fimbriatella) clathratula" = Fimbriatella3
"Mathilda (Fimbriatella) clathratula" = Fimbriatella3 snail
 Heterostropha - Pyramidellidae
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, Syrnola hoernesi3, Syrnola neumayri3, Chrysallida (Parthenina) toftlundensis3, Odostomia (Megastomia) tuexeni3, Turbonilla (Turbonilla) koeneniana3, Pyramidella (Voluspa) elata3, Pyramidella (Voluspa) plicosa3
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
Turbonilla (Turbonilla) koeneniana3 snail
sensu JANSSEN
 Heterostropha - Murchisonellidae
Ebala (Ebala) vandervlerki3
 Heterostropha - Architectonicidae
Architectonica (Architectonica) briarti3, Architectonica (Architectonica) simplex3, Architectonica (Architectonica) ariei n. sp.3, "Architectonica (Pseudotorinia) planulata" = Pseudotorinia3
Architectonica (Architectonica) ariei n. sp.3 Wienrich and Janssen 2007 sundial
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3 snail
 Heterostropha - Acteonidae
Crenilabium terebelloides3, Acteon (Acteon) semistriatus3, Acteon (Acteon) "sp. 2"3
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 2"3 Montfort 1810 snail
spelled as 'Actaeon'
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) ventricosa3, Ringicula (Ringiculina) buccinea3
 Lepetellida - Lepetellidae
Lepetella compressiuscula2
 Thecosomata - Cavoliniidae
Vaginella austriaca3
 Thecosomata - Limacinidae
Limacina miorostralis3, Limacina valvatina3
Limacina valvatina3 Reuss 1867 snail
 Opisthobranchia - Retusidae
Volvulella (Volvulella) acuminata3, Retusa sp.3, Retusa (Cylichnina) elongata3
Retusa sp.3 Brown 1827 snail
 Opisthobranchia - Cylichnidae
Bullichna paucistriata3, Scaphander grateloupi3, Cylichna (Cylichna) pseudoconvoluta3
 Opisthobranchia -
Roxania utriculus3
 Cerithioidea - Cerithiidae
Bittium (Bittium) tenuispina2, "Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
 Epitonioidea - Epitoniidae
"Amaea (Scalina) subreticula" = Scalina2, Epitonium (? Hyaloscala) "sp. 2"2, Opalia (Nodiscala) rugatina2, Opalia (Nodiscala) sp.2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
Epitonium (? Hyaloscala) "sp. 2"2 Bolten 1798 wentletrap
Opalia (Nodiscala) sp.2 De Boury 1890 wentletrap
 Sorbeoconcha - Naticidae
Naticidae indet.2, Neverita josephinia2, Polinices (Polinices) miocolligens2, Euspira gottschei2, Euspira mioaperta2, Euspira nysti2, Euspira staringi2, Euspira helicina2, Tectonatica miopusilla2, "Natica sensu lato tigrina" = Cochlis tigrina2, "Natica neglecta" = Cochlis neglecta2, Sinum (Sinum) aquense2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
Neverita josephinia2 Risso 1826 moon snail
N. josephinia olla
Euspira gottschei2 moon snail
Euspira mioaperta2 moon snail
Euspira nysti2 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
 Sorbeoconcha - Rissoidae
sensu lato Alvania antwerpiensis2, Ovirissoa gottscheana2
 Sorbeoconcha - Capulidae
Capulus ungaricus2
Capulus ungaricus2 Linnaeus 1758 snail
 Sorbeoconcha - Vitrinellidae
Circulus hennei2
Circulus hennei2 Glibert 1952 snail
 Neogastropoda - Columbellidae
Anachis (Costoanachis) hosiusi2, Anachis (Costoanachis) terebralis2, Mitrella (Macrurella) nassoides2
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2, "Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2, ? Exilifusus attenuatus2
"Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2 snail
 Neogastropoda - Nassariidae
Amyclina facki2, Nassarius bocholtensis2, Nassarius schroederi2, Nassarius sulcatus2, Nassarius cavatus2, Nassarius woodwardi2, Nassarius serraticosta2, Nassarius "sp. 2"2, "Nassarius andersoni" = Nassarius (Catilon) andersoni2, Nassarius karinae2
Nassarius "sp. 2"2 Duméril 1806 snail
"Nassarius andersoni" = Nassarius (Catilon) andersoni2 Weaver 1912 snail
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Buccinidae
Phos (Phos) decussatus2
 Neogastropoda - Cancellariidae
Trigonostoma (Trigonostoma) aperta2, Trigonostoma (Trigonostoma) banardi2, Trigonostoma (Trigonostoma) protrigonostoma2, Aneurystoma canaliculatum2, Sveltia varicosa2, Babylonella fusiformis2
Sveltia varicosa2 snail
S. varicosa paucicostata
 Neogastropoda - Mitridae
Mitra (Tiara) grateloupi3
Mitra (Tiara) grateloupi3 d'Orbigny 1852 mitre shell
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Mangeliidae
sensu lato Mangelia miorugulosa3, Benthomangelia aff. obtusangula3, Benthomangelia grippi3, Sorgenfreispira sorgenfreii3, Sorgenfreispira tenella3
Benthomangelia grippi3 Anderson 1964 snail
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Cochlespiridae
Cochlespira corneti3
 Neogastropoda - Clathurellidae
? Ceritoturris hannoverana3, Pleurotomoides borealis3, Pleurotomoides luisae3, Pleurotomoides simplex3
 Neogastropoda - Terebridae
Strioterebrum fabritzi3, Strioterebrum basteroti3, Terebra (Myurellina) acuminata3
Strioterebrum basteroti3 Nyst 1843 auger snail
Terebra (Myurellina) acuminata3 auger snail
T. acuminata neglecta
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Raphitomidae
Favriella sinuosula3, Favriella "sp. 1"3, Metuonella grippi3, Teretia anceps3, Teretia "sp. 1"3
Favriella "sp. 1"3 Hornung 1920 snail
Teretia "sp. 1"3 Norman 1888 snail
 Neogastropoda - Turridae
Boreodrillia hosiusi3, Nitidiclavus maitrejus3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula hanseata3, sensu lato Unedogemmula stoffelsii3, Gemmula (Gemmula) schafferi3, Gemmula (Gemmula) sp.3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) vermicularis3, Gemmula (Gemmula) zimmermanni3, Fusiturris aquensis3, Fusiturris duchastelii3, Acamptogenotia escheri3
Gemmula (Gemmula) sp.3 Weinkauff 1875 turrid
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
 Neogastropoda - Borsoniidae
Asthenotoma pannus3, Bathytoma jugleri3, "Genota ramosa" = Genota valeriae3, Borsonia uniplicata3
"Genota ramosa" = Genota valeriae3 Hoernes and Auinger 1891 snail
Borsonia uniplicata3 Koenen 1872 snail
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei2
Oliva (Strephona) dufresnei2 Basterot 1825 olive snail
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3, "Vexillum (Uromitra) gliberti" = Uromitra3
"Vexillum (Uromitra) gliberti" = Uromitra3 ribbed mitre
 Neogastropoda - Volutidae
Scaphella (Scaphella) bolli2, Athleta (Athleta) ficulina2
Athleta (Athleta) ficulina2 Lamarck 1811 volute
 Neogastropoda - Muricidae
Typhis (Typhis) ? pungens2, "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2, "Murex (Haustellum) inornatus" = Haustellum2, Lyrotyphis (Eotyphis) fistulosus2, Hadriania coelata2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
Hadriania coelata2 Dujardin 1837 murex snail
 Neogastropoda - Harpidae
Morum cf. dunkeri2
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Triviidae
Erato (Erato) germanica2
Erato (Erato) germanica2 Schilder 1929 snail
 Sorbeoconcha - Cassidae
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2, "Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2 Sacco 1890 snail
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Vanikoroidea - Eulimidae
Niso (Niso) terebellum2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2, Balcis (Balcis) alba2
Balcis (Balcis) alba2 snail
incl. 2specimens B. alba for. eichwaldi
Bivalvia
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Nuculanida - Yoldiidae
"Portlandia (Yoldiella) pygmaea" = Yoldiella1, Yoldia (Yoldia) glaberrima1
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
 Nuculida - Nuculidae
Nucula (Nucula) aff. nitidosa1, Nucula (Nucula) nucleus1, Nuculoma haesendoncki1
Nucula (Nucula) aff. nitidosa1 Winckworth 1930 nut clam
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
 Pectinida - Pectinoidae
Aequipecten angelonii1, Aequipecten seniensis1, Korobkovia woodi1, sensu lato Palliolum guersi n. sp.1, Pseudamussium lilli1
sensu lato Palliolum guersi n. sp.1 Wienrich 1999 scallop
Pseudamussium lilli1 Pusch 1837 scallop
 Pectinida - Anomiidae
Pododesmus (Monia) anitae1
Pododesmus (Monia) anitae1 Janse and Janssen 1983 scallop
 Arcida - Arcidae
Scapharca (Scapharca) diluvii1
 Arcida - Glycymerididae
Glycymeris (Glycymeris) obovata1
Glycymeris (Glycymeris) obovata1 Lamarck 1819 clam
G. obovata baldii
 Arcida - Limopsidae
"Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1, Limopsis (Limopsis) aurita1
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
 Cardiida - Lasaeidae
Aligena sp.1
Aligena sp.1 Lea 1843 aligena
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, Habecardium dingdense1, Parvicardium ovale1, Parvicardium straeleni1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Habecardium dingdense1 Lehmann 1885 cockle
 Cardiida - Semelidae
sensu lato Abra lehmanni1, Abra (? Abra) sorgenfreii1
 Cardiida - Tellinidae
Angulus (Laceolina) benedeni1
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Veneridae
Venus (Ventricoloidea) multilamella1, Pelecyora (Cordiopsis) polytropa1, "Gouldia minima" = Gouldia (Gouldia) minima1
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
Pelecyora (Cordiopsis) polytropa1 venus clam
P. (C.) polytropa nysti
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
 Cardiida - Mactridae
Spisula elliptica1
Spisula elliptica1 Brown 1827 clam
 Cardiida - Montacutidae
Mioerycina coarctata1
Mioerycina coarctata1 Wood 1840 clam
 Pholadida - Myidae
Sphenia pusilla1
Sphenia pusilla1 Sorgenfrel 1958 softshell clam
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Hiatellida - Hiatellidae
"Hiatella (Hiatella) arctica" = Hiatella arctica1
"Hiatella (Hiatella) arctica" = Hiatella arctica1 Linnaeus 1767 clam
H. (H.) arctica arctica
 Lucinida - Thyasiridae
Axinulus germanicus1
Axinulus germanicus1 Janssen 1972 clam
 Lucinida - Lucinidae
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1, "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1, Digitaria cf. beyschlageri1, Goodallia (Goodallia) angulata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata 274 specimens, Astarte radiata fa. convexior 56 specimens, Astarte radiata fa. goldfussi 32 specimens
Digitaria cf. beyschlageri1 clam
[entered as Digitaria beyschlagi]
Goodallia (Goodallia) angulata1 Lehmann 1885 clam
Anthozoa
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
 Scleractinia - Caryophylliidae
Ceratocyathus granulatus
Ceratocyathus granulatus Goldfuss 1826 stony coral
 Scleractinia - Turbinoliidae
"Cylindrophyllia duncani" = Peponocyathus duncani
"Cylindrophyllia duncani" = Peponocyathus duncani Reuss 1871 stony coral
 Scleractinia - Flabellidae
Flabellum sp., Flabellum cristatum
Flabellum sp. Lesson 1831 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
F. cristatum for. dietrichi
 Scleractinia - Dendrophylliidae
"Eupsammia calyculus" = Balanophyllia
"Eupsammia calyculus" = Balanophyllia stony coral
Foraminifera
 Lituolida - Lituolidae
Bigenerina nodosaria, Textularia gramen
Textularia gramen dOrbigny 1846
Globothalamea
 Rotaliida - Cibicididae
Cibicides lobatulus, Cibicides aknerianus, Cibicides dutemplei, Cibicides pseudoungerianus, Planulina austriaca
Cibicides lobatulus
incl. 4 specimens C. lobatulus var. ornatus
Cibicides dutemplei dOrbigny 1846
 Rotaliida - Nonionidae
Nonion boueanum, Nonion affine, Nonion (Porosononion) soldanii, Nonion sp.
Nonion sp. Monfort 1808
 Rotaliida - Discorbidae
Rosalina globularis, Eponides nanus
Rosalina globularis d'Orbigny 1826
Nodosariata
 Nodosariida - Nodosariidae
Vaginulina obtusicosta, Nodosaria bacillum, Nodosaria venusta, Dentalina intermittens, Dentalina inornata, Dentalina grossicosta, Dentalina acuticauda, Dentalina konincki, Lagena apiculata, Lagena striata, Lenticulina (Robulus) vortex
Dentalina konincki Reuss 1861
Lagena apiculata Reuss 1850
Lagena striata Williamson 1848
Lenticulina (Robulus) vortex Fichtel and Moll 1798
 Polymorphinida - Polymorphinidae
Pseudopolymorphina incerta, Pseudopolymorphina ovalis, Globulina gibba, Guttulina austriaca, Guttulina problema, Sigmomorphina sp.
Globulina gibba d'Orbigny 1832
incl. 1 specimen G. gibba for. tubulosa
Guttulina problema d'Orbigny 1826
Sigmomorphina sp. Cushman and Ozawa 1928
 Vaginulinida - Vaginulinidae
Lenticulina (Vaginulinopsis) gerlachi, Lenticulina (Vaginulinopsis) hauerina, Lenticulina (Robulus) grandis, Lenticulina (Robulus) ornata, Lenticulina (Planularia) grundensis, Lenticulina (Planularia) kubinyi, Lenticulina (Lenticulina) inornata
Foraminifera
 Nodosariacea - Glandulinidae
Glandulina ovula
 Asterigerinacea - Asterigerinidae
Asterigerina guerichi
Asterigerina guerichi
A. gürichi staeschi
 Discorbacea - Bagginidae
Cancris auriculus
Cancris auriculus Fichtel and Moll 1798
 Buliminacea - Uvigerinidae
Uvigerina semiornata, Uvigerina tenuipustulata
Uvigerina semiornata
U. semiornata for. semiornata
Tubothalamea
 Miliolida - Miliolidae
Miliolidae indet., Sigmoilina tenuis, Pyrgo bulloides
Miliolidae indet. Ehrenberg 1839
 Miliolida - Hauerinidae
Triloculina inflata, Triloculina scapha
Foraminifera
 Rotalioidea - Elphidiidae
Elphidium inflatum, Elphidium ungeri
 Lituolacea - Ataxophragmiidae
Martinottiella communis, Karreriella siphonella
Martinottiella communis Huang et al. 2019
  - Spiroplectamminidae
Spiroplectammina carinata
 Buliminoidea - Buliminidae
Virgulina (Virgulinella) pertusa
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007