Bore Lullingen 1, depth 33m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61903: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Rhynchonellata
 Rhynchonellida - Cryptoporidae
Cryptopora sp.1
Cryptopora sp.1 Jeffreys 1869
 Terebratulida - Terebratulidae
Terebratula sp.1
Terebratula sp.1 Müller 1776
Bryozoa
  -
? Bryozoa indet.1
? Bryozoa indet.1 Ehrenberg 1831
Gymnolaemata
 Cheilostomata - Membraniporidae
Reussirella haidingeri1, Discoporella umbellata1
Discoporella umbellata1 Defrance 1823
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
Cephalopoda
 Spirulida - Spirulirostridae
Spirulirostra hoernesi1
Scaphopoda
 Dentaliida - Dentaliidae
Lentigodentalium pseudomutabile1, "Pseudantalis aturensis" = Dentalium (Pseudantilis)1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
 Gadilida - Gadilidae
Gadila dingdenensis1
Gastropoda
 Heterostropha - Mathildidae
Gegania cancellata3, sensu lato Mathilda concinna3
 Heterostropha - Pyramidellidae
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, "Turbonilla (Pyrgolampros) "sp. 6"" = Pyrgolampros3, Syrnola dubia3, Syrnola hoernesi3, Syrnola neumayri3, Syrnola "sp. 1"3, Chrysallida (Pyrgulina) pygmaea3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Megastomia) cf. conoidea3, Odostomia (Megastomia) cf. pallidaeformis3, Odostomia (Megastomia) tuexeni3, Turbonilla (Mormula) "sp. 3"3, Turbonilla (Mormula) "sp. 4"3, Turbonilla (Turbonilla) gastaldi3, Turbonilla (Turbonilla) koeneniana3, Turbonilla (Turbonilla) "sp. 2"3, Pyramidella (Voluspa) elata3, Pyramidella (Voluspa) plicosa3
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) "sp. 6"" = Pyrgolampros3 Sacco 1892 snail
Syrnola "sp. 1"3 Adams 1860 snail
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
Ch. pygmaea f. belgica
Turbonilla (Mormula) "sp. 3"3 Adams 1864 snail
Turbonilla (Mormula) "sp. 4"3 Adams 1864 snail
Turbonilla (Turbonilla) gastaldi3 snail
sensu JANSSEN
Turbonilla (Turbonilla) koeneniana3 snail
sensu JANSSEN
 Heterostropha - Acteonidae
Acteon (Acteon) arnumensis3, Acteon (Acteon) laevigatus3, Acteon (Acteon) semistriatus3
Acteon (Acteon) arnumensis3 Sorgenfrei 1958 snail
spelled as 'Actaeon'
Acteon (Acteon) laevigatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) ? laureatum2
 Opisthobranchia - Bullidae
"Tornatina bellardi" = Bulla (Tornatina)3
"Tornatina bellardi" = Bulla (Tornatina)3 snail
 Opisthobranchia - Retusidae
Volvulella (Volvulella) acuminata3
 Opisthobranchia - Cylichnidae
Bullichna paucistriata3, Scaphander grateloupi3, Cylichna (Cylichna) pseudoconvoluta3
 Opisthobranchia -
Roxania utriculus3
 Cerithioidea - Cerithiidae
Bittium (Bittium) tenuispina2, "Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2
 Epitonioidea - Epitoniidae
"Amaea (Scalina) subreticula" = Scalina2, Epitonium (Spiniscala) frondiculum2, Opalia (Pliciscala) pertusa2, Opalia (Nodiscala) rugatina2, sensu lato Cirsotrema "sp. 1"2, Cirsotrema (Cirsotrema) crassicostatum2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
Epitonium (Spiniscala) frondiculum2 Wood 1842 wentletrap
sensu lato Cirsotrema "sp. 1"2 Mörch 1852 wentletrap
Cirsotrema (Cirsotrema) crassicostatum2 Deshayes 1839 wentletrap
 Sorbeoconcha - Naticidae
Naticidae indet.2, Neverita josephinia2, Polinices (Polinices) miocolligens2, Euspira edithae2, Euspira gottschei2, Euspira staringi2, Euspira helicina2, Tectonatica miopusilla2, "Natica sensu lato tigrina" = Cochlis tigrina2, "Natica neglecta" = Cochlis neglecta2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
Neverita josephinia2 Risso 1826 moon snail
N. josephinia olla
Euspira edithae2 moon snail
Euspira gottschei2 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
 Sorbeoconcha - Triphoridae
Norephora (Norephora) fritschi2
 Sorbeoconcha - Triforidae
Cerithiella (Cerithiella) genei2
 Sorbeoconcha - Rissoidae
Alvania (Alvania) partschi2, Alvania (Alvania) basisulcata2, Ovirissoa gottscheana2
 Sorbeoconcha - Tornidae
Tornus (Tornus) trigonostoma2
 Sorbeoconcha - Vitrinellidae
Circulus praecedens2, Circulus partimstriatus2, Circulus hennei2
Circulus praecedens2 snail
12 specimen C. praecedens for. praecedens and 1 specimen C. praecedens for. gliberti
Circulus hennei2 Glibert 1952 snail
 Neogastropoda - Columbellidae
Anachis (Costoanachis) hosiusi2, Mitrella (Macrurella) nassoides2
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius karinae2
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Cancellariidae
Trigonostoma (Trigonostoma) aperta2, Trigonostoma (Trigonostoma) extractrix2, Cancellaria (Merica) contorta2, Cancellaria (Bivetiella) cancellata2, Sveltia varicosa2, Babylonella fusiformis2
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
Cancellaria (Bivetiella) cancellata2 Linnaeus 1767 snail
C. cancellata praecedens
Sveltia varicosa2 snail
S. varicosa paucicostata
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Mangeliidae
Benthomangelia aff. obtusangula3, Sorgenfreispira tenella3
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3, Pleurotomoides simplex3
 Neogastropoda - Terebridae
Strioterebrum hoernesi3, Strioterebrum ronaldi3, Terebra (Myurellina) acuminata3
Strioterebrum ronaldi3 auger snail
Terebra (Myurellina) acuminata3 auger snail
T. acuminata neglecta
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Raphitomidae
Favriella sinuosula3, Mioawateria andersoni3, Metuonella grippi3, Teretia anceps3, Pleurotomella "sp. 1"3
Pleurotomella "sp. 1"3 Verrill 1872 snail
 Neogastropoda - Turridae
Boreodrillia hosiusi3, Nitidiclavus maitrejus3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula hanseata3, sensu lato Unedogemmula stoffelsii3, Gemmula (Gemmula) sp.3, Gemmula (Gemmula) zimmermanni3, Fusiturris aquensis3, Fusiturris duchastelii3, Ingaunoturricula marginata3
Gemmula (Gemmula) sp.3 Weinkauff 1875 turrid
Gemmula n. sp.
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
 Neogastropoda - Borsoniidae
Asthenotoma pannoides3, Asthenotoma pannus3, Asthenotoma festiva3, Bathytoma jugleri3, "Genota ramosa" = Genota valeriae3, Borsonia uniplicata3
"Genota ramosa" = Genota valeriae3 Hoernes and Auinger 1891 snail
Borsonia uniplicata3 Koenen 1872 snail
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei2
Oliva (Strephona) dufresnei2 Basterot 1825 olive snail
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3
 Neogastropoda - Volutidae
Scaphella (Scaphella) bolli2
 Neogastropoda - Muricidae
Typhis (Typhis) ? pungens2, "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2, Coralliophila (Aldrichia) cf. granifera2, Pterynotus (Alipurpura) parvifolius2, Lyrotyphis (Eotyphis) fistulosus2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Cassidae
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2, "Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2 Sacco 1890 snail
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Vanikoroidea - Eulimidae
Niso (Niso) terebellum2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2, Balcis (Balcis) alba2
Bivalvia
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1, Nuculana (Jupiteria) curvirostris1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
Nuculana (Jupiteria) curvirostris1 de Cserna 1961 pointed nut clam
 Nuculanida - Yoldiidae
"Portlandia (Yoldiella) pygmaea" = Yoldiella1, Yoldia (Yoldia) glaberrima1
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
incl. 1specimen of Portlandia pygm. fa. alta
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
 Nuculida - Nuculidae
Nucula (Nucula) nucleus1, Nuculoma haesendoncki1
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
 Pectinida - Pectinoidae
Aequipecten angelonii1, Korobkovia woodi1, Pseudamussium lilli1
Pseudamussium lilli1 Pusch 1837 scallop
 Pectinida - Limidae
Limatula sulcata1
Limatula sulcata1 file clam
 Pectinida - Anomiidae
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Ostreida - Ostreidae
Cubitostrea digitalina1, ? Ostrea cf. edulis1
Cubitostrea digitalina1 Eichwald 1830 oyster
? Ostrea cf. edulis1 Linnaeus 1758 oyster
 Arcida - Arcidae
Bathyarca pectunculoides1, Scapharca (Scapharca) diluvii1
Bathyarca pectunculoides1 Scacchi 1834 clam
Scapharca (Scapharca) diluvii1 Lamarck 1805 clam
 Arcida - Limopsidae
"Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1, Limopsis (Limopsis) aurita1
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
 Cardiida - Cardiidae
Papillicardium papillosum1, "Habecardium subturgidum" = Laevicardium (Habecardium)1, Parvicardium straeleni1
Papillicardium papillosum1 Poli 1781 cockle
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
 Cardiida - Semelidae
Abra (Abra) antwerpiensis1, Abra (? Abra) sorgenfreii1
Abra (Abra) antwerpiensis1 Glibert 1945 abra
 Cardiida - Solecurtidae
Solecurtus basteroti1, Azorinus (Azorinus) chamasolen1
Azorinus (Azorinus) chamasolen1 clam
A. chamasolen miocaenicus
 Cardiida - Tellinidae
Angulus (Laceolina) benedeni1
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Veneridae
Venus (Ventricoloidea) multilamella1, Callista chione1, "Gouldia minima" = Gouldia (Gouldia) minima1
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
Callista chione1 Linnaeus 1758 venus clam
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
 Cardiida - Mactridae
Lutraria frangula1
 Cardiida - Montacutidae
Montacuta cf. caecilia1
 Cardiida - Erycinidae
Erycina aff. backlundi1
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Pholadida - Teredinidae
Teredinidae indet.1
Teredinidae indet.1 Rafinesque 1815 clam
 Solenida - Pharidae
Cultellus (Cultellus) aff. cultellatus1
 Thraciida - Thraciidae
Thracia (Thracia) aff. inflata1
 Lucinida - Lucinidae
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1, Anodontia (Loripinus) noerregaardi1
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata
Anthozoa
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
 Scleractinia - Flabellidae
Flabellum sp., Flabellum cristatum, Flabellum tuberculatum
Flabellum sp. Lesson 1831 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
F. cristatum for. dietrichi
Flabellum tuberculatum Keferstein 1859 stony coral
Foraminifera
 Lituolida - Lituolidae
Textularia gramen
Textularia gramen dOrbigny 1846
Globothalamea
 Rotaliida - Cibicididae
Cibicides lobatulus, Cibicides dutemplei, Cibicides pseudoungerianus
Cibicides lobatulus
incl. 1 specimen C. lob. var. ornatus
Cibicides dutemplei dOrbigny 1846
 Rotaliida - Gavelinellidae
Gyroidina soldanii
Nodosariata
 Nodosariida - Nodosariidae
Vaginulina legumen, Nodosaria bacillum, Dentalina konincki
Dentalina konincki Reuss 1861
 Polymorphinida - Polymorphinidae
Pseudopolymorphina subnodosa, Globulina gibba, Guttulina problema
Globulina gibba d'Orbigny 1832
Guttulina problema d'Orbigny 1826
 Vaginulinida - Vaginulinidae
Lenticulina (Robulus) cultrata, Lenticulina (Robulus) orbicularis, Lenticulina (Robulus) cassis, Lenticulina (Planularia) grundensis, Lenticulina (Lenticulina) inornata, Lenticulina (Lenticulina) calcar
Tubothalamea
 Miliolida - Miliolidae
Sigmoilina tenuis
Foraminifera
 Rotalioidea - Elphidiidae
Elphidium inflatum
  - Spiroplectamminidae
Spiroplectammina carinata
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007