Bore Lullingen 1, depth 39m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61896: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Rhynchonellata
 Rhynchonellida - Cryptoporidae
Cryptopora sp.1
Cryptopora sp.1 Jeffreys 1869
Gymnolaemata
 Cheilostomata - Membraniporidae
Reussirella haidingeri1, Discoporella umbellata1, Biflustra savartii1
Discoporella umbellata1 Defrance 1823
Biflustra savartii1 Audouin 1826
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
 Cheilostomata - Cellariidae
Cellaria marginata1
Polychaeta
  -
Polychaeta indet.1
Polychaeta indet.1 Grube 1850
Errantida indet. A
 Sabellida - Serpulidae
Serpulidae indet.1, Ditrupa cornea1
Serpulidae indet.1 Rafinesque 1835
Serpulidae indet. B
Scaphopoda
 Dentaliida - Dentaliidae
Lentigodentalium pseudomutabile1, Fissidentalium twistringense1, "Pseudantalis aturensis" = Dentalium (Pseudantilis)1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
 Gadilida - Gadilidae
Gadila dingdenensis1
Gastropoda
 Heterostropha - Pyramidellidae
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, Syrnola hoernesi3, Syrnola "sp. 2"3, Chrysallida (Pyrgulina) "sp. 2"3, Chrysallida (Pyrgulina) pygmaea3, Chrysallida (Parthenina) cimbrica3, Chrysallida (Parthenina) modesta3, Chrysallida (Parthenina) "sp. 1"3, Chrysallida (Parthenina) "sp. 3"3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Megastomia) cf. conoidea3, Odostomia (Megastomia) tuexeni3, Odostomia (Odostomia) sp.3, Evalea sp.3, Turbonilla (Mormula) "sp. 4"3, Turbonilla (Turbonilla) gastaldi3, Turbonilla (Turbonilla) koeneniana3, Eulimella (Eulimella) acicula3, Eulimella (Eulimella) scillae3
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
Syrnola "sp. 2"3 Adams 1860 snail
Chrysallida (Pyrgulina) "sp. 2"3 Carpenter 1856 snail
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
incl. 2 specimens Chrysallida pygmaea f. belgica
Chrysallida (Parthenina) "sp. 1"3 Bucquoy et al. 1883 snail
Chrysallida (Parthenina) "sp. 3"3 Bucquoy et al. 1883 snail
Odostomia (Odostomia) sp.3 Fleming 1817 snail
Evalea sp.3 Adams 1860 snail
Turbonilla (Mormula) "sp. 4"3 Adams 1864 snail
 Heterostropha - Architectonicidae
Architectonica (Architectonica) briarti3, "Architectonica (Pseudotorinia) planulata" = Pseudotorinia3
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3 snail
 Heterostropha - Acteonidae
Acteon (Acteon) laevigatus3, Acteon (Acteon) "sp. 1"3, Acteon (Acteon) "sp. 2"3
Acteon (Acteon) laevigatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 1"3 Montfort 1810 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 2"3 Montfort 1810 snail
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Lepetellida - Lepetellidae
Lepetella compressiuscala2
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) ? laureatum2
 Trochoidea - Skeneidae
Skenea minuta2
 Thecosomata - Limacinidae
Limacina miorostralis3
 Opisthobranchia - Retusidae
Volvulella (Volvulella) acuminata3, Retusa (Cylichnina) elongata3
 Opisthobranchia - Cylichnidae
Scaphander grateloupi3, Cylichna (Cylichna) pseudoconvoluta3
 Opisthobranchia -
Roxania utriculus3
 Cocculinoidea - Cocculinidae
Cocculina miocaenica2
 Cerithioidea - Cerithiidae
"Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
 Epitonioidea - Epitoniidae
"Amaea (Scalina) subreticula" = Scalina2, Epitonium (Spiniscala) frondiculum2, Opalia (Pliciscala) pertusa2, Opalia (Nodiscala) rugatina2, sensu lato Cirsotrema ertborni2, Cirsotrema (Cirsotrema) crassicostatum2, Acirsa (Plesioacirsa) mioplicatula2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
Epitonium (Spiniscala) frondiculum2 Wood 1842 wentletrap
Cirsotrema (Cirsotrema) crassicostatum2 Deshayes 1839 wentletrap
 Sorbeoconcha - Naticidae
Naticidae indet.2, Euspira staringi2, Euspira helicina2, Tectonatica miopusilla2, "Natica sensu lato tigrina" = Cochlis tigrina2, "Natica neglecta" = Cochlis neglecta2, Sinum (Sinum) aquense2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
 Sorbeoconcha - Cerithiopsidae
Cerithiopsis (Cerithiopsis) vogeli2
 Sorbeoconcha - Triphoridae
Norephora (Norephora) fritschi2
 Sorbeoconcha - Rissoidae
sensu lato Alvania holsatica2, Ovirissoa gottscheana2, Ovirissoa westfalica2
 Sorbeoconcha - Tornidae
Tornus (Tornus) trigonostoma2
 Sorbeoconcha - Vitrinellidae
Circulus praecedens2, Circulus partimstriatus2, Circulus hennei2
Circulus praecedens2 snail
C. praecedens for. praecedens
Circulus hennei2 Glibert 1952 snail
 Neogastropoda - Columbellidae
Anachis (Costoanachis) "sp. 1"2, Mitrella (Macrurella) nassoides2
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2, ? Exilifusus attenuatus2
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius voorthuyseni2, Nassarius karinae2
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Buccinidae
Phos (Phos) decussatus2
 Neogastropoda - Cancellariidae
Trigonostoma (Trigonostoma) extractrix2, Cancellaria (Merica) bellardi2, Cancellaria (Merica) contorta2, Sveltia varicosa2, Babylonella fusiformis2
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
Sveltia varicosa2 snail
S. varicosa paucicostata
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Mangeliidae
sensu lato Mangelia andersoni3, Benthomangelia aff. obtusangula3, Sorgenfreispira sorgenfreii3, Sorgenfreispira tenella3
sensu lato Mangelia andersoni3 Wienrich and Janssen 2007 snail
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3, Pleurotomoides simplex3
 Neogastropoda - Terebridae
Strioterebrum hoernesi3, Strioterebrum ronaldi3, Terebra (Myurellina) acuminata3
Strioterebrum ronaldi3 auger snail
Terebra (Myurellina) acuminata3 auger snail
T. acuminata neglecta
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Raphitomidae
Favriella sinuosula3, Metuonella grippi3, Teretia anceps3, sensu lato Teretia "sp. 2"3, Raphitoma aff. praehispida3
sensu lato Teretia "sp. 2"3 Norman 1888 snail
 Neogastropoda - Turridae
Perrona obliquiplicatula3, Boreodrillia hosiusi3, Nitidiclavus maitrejus3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula stoffelsii3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) zimmermanni3, Fusiturris aquensis3, Fusiturris duchastelii3
sensu lato Unedogemmula boreoturricula3 turrid
incl. 8 specimens U. (s. lat.) boreoturricula x hanseata ?
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
 Neogastropoda - Borsoniidae
Asthenotoma pannoides3, Asthenotoma pannus3, Bathytoma jugleri3, "Genota ramosa" = Genota valeriae3
"Genota ramosa" = Genota valeriae3 Hoernes and Auinger 1891 snail
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3, "Vexillum (Uromitra) gliberti" = Uromitra3
"Vexillum (Uromitra) gliberti" = Uromitra3 ribbed mitre
 Neogastropoda - Volutidae
Scaphella (Scaphella) bolli2
 Neogastropoda - Muricidae
Typhis (Typhis) ? pungens2, "Murex (Haustellum) inornatus" = Haustellum2, Coralliophila (Aldrichia) cf. granifera2, Lyrotyphis (Eotyphis) fistulosus2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Triviidae
Erato (Erato) germanica2
Erato (Erato) germanica2 Schilder 1929 snail
 Sorbeoconcha - Cassidae
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2, "Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2 Sacco 1890 snail
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Vanikoroidea - Eulimidae
Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2, Balcis (Balcis) alba2
Balcis (Balcis) alba2 snail
incl. 2 specimens B. alba for. lactea
Bivalvia
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Nuculanida - Yoldiidae
"Portlandia (Yoldiella) pygmaea" = Yoldiella1, Yoldia (Yoldia) glaberrima1
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
 Nuculida - Nuculidae
Nucula (Lamellinucula) jeffreysi1, Nucula (Nucula) nucleus1, Nuculoma haesendoncki1
Nucula (Lamellinucula) jeffreysi1 Bellardi 1875 nut clam
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
 Mytilida - Crenellidae
Musculus (Musculus) cf. sternbergensis1
 Mytilida - Mytilidae
Amygdalum aff. longum1
 Pectinida - Pectinoidae
Aequipecten angelonii1, Korobkovia woodi1, Pseudamussium lilli1
Pseudamussium lilli1 Pusch 1837 scallop
 Pectinida - Limidae
Limea (Limea) aff. strigilata1
Limea (Limea) aff. strigilata1 Brocchi 1814 file clam
 Pectinida - Anomiidae
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Ostreida - Ostreidae
Ostreidae indet.1
Ostreidae indet.1 Rafinesque 1815 oyster
 Arcida - Arcidae
Bathyarca pectunculoides1, Scapharca (Scapharca) diluvii1
Bathyarca pectunculoides1 Scacchi 1834 clam
Scapharca (Scapharca) diluvii1 Lamarck 1805 clam
 Arcida - Limopsidae
"Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1, Limopsis (Limopsis) aurita1
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
 Cardiida - Lasaeidae
"Mysella (Mysella) bidentata" = Diarmaidia bidentata1
"Mysella (Mysella) bidentata" = Diarmaidia bidentata1 Montague 1803 clam
 Cardiida - Leptonidae
Lepton (? Lepton) transversarium1, Properycina (Properycina) edlaueri1
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, Habecardium dingdense1, Parvicardium straeleni1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Habecardium dingdense1 Lehmann 1885 cockle
 Cardiida - Semelidae
sensu lato Abra lehmanni1, Abra (Abra) antwerpiensis1, Abra (? Abra) sorgenfreii1
Abra (Abra) antwerpiensis1 Glibert 1945 abra
 Cardiida - Solecurtidae
Solecurtus basteroti1
 Cardiida - Tellinidae
"Macoma elliptica" = Laciolina magna1, Angulus (Laceolina) benedeni1
"Macoma elliptica" = Laciolina magna1 Spengler 1798 tellin clam
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Psammobiidae
Gari (Psammotaena) aff. angusta1
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Veneridae
Venus (Ventricoloidea) multilamella1, Pelecyora (Cordiopsis) polytropa1, Callista chione1, "Gouldia minima" = Gouldia (Gouldia) minima1
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
Pelecyora (Cordiopsis) polytropa1 venus clam
P. (C.) polytropa nysti
Callista chione1 Linnaeus 1758 venus clam
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
 Cardiida - Mactridae
Lutraria frangula1
 Pholadida - Myidae
Sphenia pusilla1
Sphenia pusilla1 Sorgenfrel 1958 softshell clam
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Pholadida - Teredinidae
Teredinidae indet.1
Teredinidae indet.1 Rafinesque 1815 clam
 Solenida - Pharidae
Cultellus (Cultellus) aff. cultellatus1, Ensis hausmanni1
 Poromyida - Cuspidariidae
Cuspidaria sp.1
Cuspidaria sp.1 Dall 1886 clam
 Thraciida - Thraciidae
Thracia (Thracia) aff. inflata1
 Thraciida - Periplomatidae
Periplomatidae indet.1
Periplomatidae indet.1 Dall 1895 clam
 Lucinida - Lucinidae
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1, "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
 Carditida - Condylocardiidae
Erycinella chavani1
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1, Goodallia (Goodallia) angulata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata 27 specimens, Astarte radiata fa. convexior 5 specimens
Goodallia (Goodallia) angulata1 Lehmann 1885 clam
Anthozoa
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
 Scleractinia - Flabellidae
Flabellum sp., Flabellum cristatum
Flabellum sp. Lesson 1831 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
Foraminifera
 Lituolida - Lituolidae
Textularia gramen
Textularia gramen dOrbigny 1846
Globothalamea
 Rotaliida - Cibicididae
Cibicides lobatulus, Cibicides pseudoungerianus
Nodosariata
 Nodosariida - Nodosariidae
Dentalina "sp. 1", Dentalina konincki
Dentalina "sp. 1" d'Orbigny 1826
Dentalina konincki Reuss 1861
 Polymorphinida - Polymorphinidae
Pseudopolymorphina subnodosa, Globulina gibba, Guttulina austriaca, Guttulina problema, Sigmomorphina sp.
Pseudopolymorphina subnodosa
incl. 1 specimen P. subn. for. rugosa
Globulina gibba d'Orbigny 1832
Guttulina problema d'Orbigny 1826
Sigmomorphina sp. Cushman and Ozawa 1928
 Vaginulinida - Vaginulinidae
Lenticulina (Vaginulinopsis) hauerina, Lenticulina (Robulus) cultrata, Lenticulina (Lenticulina) calcar, Lenticulina (Lenticulina) gibba
Foraminifera
 Rotalioidea - Elphidiidae
Elphidium inflatum
 Lituolacea - Ataxophragmiidae
Martinottiella communis
Martinottiella communis Huang et al. 2019
  - Spiroplectamminidae
Spiroplectammina carinata
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007