Portezuelo Sumich Norte, Pinturas Fm., Lago Buenos Aires (Miocene of Argentina)

Where: Santa Cruz, Argentina (47.0° S, 70.6° W: paleocoordinates 47.7° S, 64.9° W)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Pinturan zone, Pinturas Formation, Burdigalian (20.4 - 16.0 Ma)

• Pertains to "Middle Sequence" = "Pinturan" of Kramarz & Bellosi 2005

•Brandoni et al 2016: The absolute ages of ca. 17.99 Ma near the base of the Pinturas Formation at Estancia el Carmen (47? 120 44.6400S, 70? 340 9700W), and ca. 16.8 Ma in the Santa Cruz For- mation that overlies the Pinturas Formation at Portezuelo Sumich Norte (see Perkins et al. 2012 and Fleagle et al. 2012) (Fig. 1b) indicate that the lower and middle sequences from the Pinturas Formation would be similar in age to those from the coastal localities of the Santa Cruz Formation (‘‘typical Santacrucian fauna’’) with ages of ca. 18–16 Ma (Bown and Fleagle 1993; Fleagle et al. 1995; Tejedor et al. 2006; Perkins et al. 2012), and younger than those from the lower Fossil Zone of the Colhue-Huapi Member of the Sarmiento Formation (‘‘Colhuehuapian fauna’’) (Kramarz et al. 2010) with ages of ca. 20.4–20.0 Ma (Re´ et al. 2010) or 21.11–18.62 Ma (Dunn et al. 2013) (Fig. 2)

•Brandoni et al 2016: In the locality of Portezuelo Sumich Norte (46? 570

•33.0900S, 70? 400 200W; Fig. 1a, b), numerous fossil remains (including primates) come from opposite sides of a badland ridge that probably preserves a part of the base of the middle sequence that has filled a scour cur into the top of the lower sequence (Bown and Larriestra 1990 fig. 9a; T. Bown, Pers. Com). There is an absolute age of ca. 16.8 Ma from a tuff of the Santa Cruz Formation that lies uncon- formably over the top of the Pinturas sequence (Bown and Larriestra 1990; Perkins et al. 2012; Fleagle et al. 2012);

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; volcaniclastic sediments

• Brandoni et al 2016: The Pinturas Formation is composed of up to 100 m of Miocene continental pyroclastic and eolian epiclastic sed-iments outcropping in the upper valley of the Pinturas River and its tributaries (Fig. 1a). It lies unconformably atop the Jurassic La Matilde Formation (Bahı´a Laura River and its tributaries (Fig. 1a). It lies unconformably atop the Jurassic La Matilde Formation (Bahı´a Laura Group), and is divided by two intraformational erosional unconformities into three sequences: (a) the lower sequence, dominated by pyroclastic mudrocks upon which very mature paleosols were formed; (b) the middle sequence, composed largely by epiclastic sand occurring as barchanoid paleodunes; and (c) the upper sequence, con-sisting of massive, poorly bedded pyroclastic mudrocks (Bown et al. 1988; Bown and Larriestra 1990). The Pin-turas Formation differs from the Santa Cruz Formation in the greater volume of pyroclastic materials, absence of marine intercalations, and the presence of mature paleosols in the lower sequence (Bown and Larriestra 1990).

Size class: macrofossils

Reposited in the MACN

Primary reference: T. M. Bown and C. N. Larriestra. 1990. Sedimentary paleoenvironments of fossil platyrrhine localities, Miocene Pinturas Formation, Santa Cruz Province, Argentina. Journal of Human Evolution 19(1-2):87-119 [D. Croft/D. Croft]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 59319: authorized by Darin Croft, entered by Darin Croft on 26.03.2006, edited by Juan Carrillo, Jelle Zijlstra, Miranta Kouvari, Grace Varnham and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

Mammalia
 Primates -
Platyrrhini indet.13 Geoffroy 1812 monkey
MACN-PV SC35, P2?
 Primates - Homunculidae
Soriacebus sp.15 Fleagle et al. 1987 monkey
2 specimens
Soriacebus cf. ameghinorum16 Fleagle et al. 1987 monkey
1 specimen
cf. Soriacebus ameghinorum13 Fleagle et al. 1987 monkey
MPM-PV 17395, right lower canine; MPM PV 17412, 17396, 17415
Soriacebus ameghinorum13 Fleagle et al. 1987 monkey
MPM-PV 17394, maxilla P3-4; MPM PV 17413
 Primates -
cf. Carlocebus carmenensis15 Fleagle 1990 monkey
2 specimens
Carlocebus carmenensis7 Fleagle 1990 monkey
7 specimens
Carlocebus intermedius n. sp.7 Fleagle 1990 monkey
Holotype plus 1 specimen
 Primates - Pitheciidae
Pitheciidae indet.14 Mivart 1865 monkey
MPM PV 17414
 Rodentia -
Eosteiromys annectens9 Ameghino 1901 caviomorph
5 specimens
Eosteiromys cf. homogenidens9 Ameghino 1902 caviomorph
2 specimens
 Rodentia - Erethizontidae
Branisamyopsis praesigmoides9 Kramarz 2004 porcupine
4 specimens
 Rodentia -
Steiromys principalis9 Ameghino 1901 caviomorph
1 specimen
Luantus propheticus10 Ameghino 1899 caviomorph
13 specimens
Neoreomys pinturensis n. sp.11 Kramarz 2006 caviomorph
Neoreomys australis11 Ameghino 1887 caviomorph
1 specimen
 Rodentia - Dinomyidae
Scleromys quadrangulatus11 Kramarz 2006 caviomorph
9 specimens
 Rodentia - Neoepiblemidae
Perimys intermedius8 Kramarz 2002 caviomorph
7 specimens
 Rodentia -
Acarechimys sp.9 Patterson 1965 caviomorph
1 specimen; probably a new species, but material is too fragmentary for definitive diagnosis
Acarechimys minutissimus1 Ameghino 1887 caviomorph
MACN-PV SC2158
 Rodentia - Echimyidae
Prostichomys bowni9 Kramarz 2001 spiny rat
31 specimens
 Rodentia - Acaremyidae
Galileomys eurygnathus9 Kramarz 2004 caviomorph
14 specimens
 Panameriungulata - Proterotheriidae
Tetramerorhinus prosistens12 Ameghino 1899 placental
8 specimens
Tetramerorhinus fleaglei n. sp.12 Kramarz and Bond 2005 placental
holotype plus 10 specimens
Lambdaconus lacerum12 Ameghino 1902 placental
2 specimens
Picturotherium migueli12 Kramarz and Bond 2005 placental
2 specimens
 Megatherioidea - Megalonychidae
Hapalops sp.4 Ameghino 1887 edentate
MPM-PV 17409
  - Mylodontidae
Nematherium sp.5 Ameghino 1887 edentate
MPM-PV 17422
 Microbiotheria - Microbiotheriidae
"Microbiotherium divisum" = Oligobiotherium divisum6 Ameghino 1902 marsupial
MACN Pv SC520, dentary m2-3
Pachybiotherium illuminatum2 Goin et al. 2010 marsupial
MACN-PV SC3617b
 Paucituberculata - Palaeothentidae
Acdestis oweni3 Ameghino 1887 marsupial
MACN SC numbers 407, 408, 416, 451, 476, 500