PBDB Collection

UCMP V6168, Avenay quarry (Eocene of France)

Also known as Avenay-Val-d’Or

Where: Marne, France (49.1° N, 4.0° E: paleocoordinates 44.5° N, 1.3° E)

• coordinate based on unpublished field data

• small collection-level geographic resolution

When: Lignites de Soissonais Formation, Ypresian (56.0 - 47.8 Ma)

• late Sparnacian, NP12, MP8+9 (Smith & Smith, 2003)

• formation-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size class: mesofossils

Primary reference: D. E. Russell, M. Godinot, P. Louis and D. E. Savage. 1979. Apatotheria (Mammalia) de l'Eocène inférieur de France et de Belgique. Bulletin du Muséum National d'Histoire Naturelle, section C: Science de la Terre 1(3):203-243 [M. Carrano/M. Carrano/J. Alroy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 35535: authorized by Matthew Carrano, entered by Matthew Carrano on 19.12.2003, edited by Philip Mannion, John Alroy, K. Christopher Beard, Jason Head, Torsten Liebrecht, Terri Cleary, David Nicholson, Patricia Holroyd and Richard Butler

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Reptilia
Squamata - Gekkonidae	Gekkonidae indet.² Gekkonidae indet.² Gray 1825 gecko fragment of left dentary
Squamata - Iguanidae	"Geiseltaliellus louisi" = Geiseltaliellus longicaudus¹ "Geiseltaliellus louisi" = Geiseltaliellus longicaudus¹ Kuhn 1944 squamates 1 right dentary (n AV 16462)
Squamata - Agamidae	Tinosaurus sp.¹, Tinosaurus europeocaenus² Tinosaurus sp.¹ Marsh 1872 squamates 1 left dentary (n CB 16437) Tinosaurus europeocaenus² Auge and Smith 1997 squamates MHNH AV 16464 ,anterior left dentary
Squamata - Boidae	Calamagras gallicus¹⁰ Calamagras gallicus¹⁰ Rage 1977 dwarf boa Rage (1977) mentions a total of 30 referred vertebrae which come from Avenay and other localities in the Paris Basin, but does not give details on specimen and catalogue numbers for individual localities
Squamata - Tropidophiidae	Dunnophis matronensis⁹ Dunnophis matronensis⁹ Rage 1973 dwarf boa Rage (1973) mentions "nombreuses vertèbres" which come from Avenay and other localities in the Paris Basin, but does not give details on specimen and catalogue numbers for individual localities
Squamata - Lacertidae	Dormaalisaurus girardoti² Dormaalisaurus girardoti² Augé and Smith 2002 squamates incomplete left and right dentaries
Squamata - Eolacertidae	Eolacerta sp.² Eolacerta sp.² Nöth 1940 squamates dorsal vertebra
Squamata - Necrosauridae	Necrosaurus sp.² Necrosaurus sp.² Filhol 1876 squamates MNHN, three osteoderms
Squamata - Amphisbaenidae	Amphisbaenidae indet.² Amphisbaenidae indet.² Gray 1865 worm lizard Av 7927, a vertebra; Av 7923, fragment of dentary
Testudines - Carettochelyidae	Allaeochelys sp.³ Allaeochelys sp.³ Noulet 1867 turtle
Testudines - Geoemydidae	Palaeochelys sp.³ Palaeochelys sp.³ Meyer 1847 turtle
Testudines - Trionychidae	Trionyx sp.³, "Palaeotrionyx vittatus" = Axestemys vittata³ Trionyx sp.³ Geoffroy Saint-Hilaire 1809 softshell turtle "Palaeotrionyx vittatus" = Axestemys vittata³ Pomel 1847 softshell turtle
Testudines - Podocnemididae	Neochelys eocaenica⁷ Neochelys eocaenica⁷ de Stefano 1902 sideneck turtle
Mammalia
Macroscelidea - Amphilemuridae	Macrocranion cf. nitens⁵ Macrocranion cf. nitens⁵ Matthew 1918 elephant shrew Russell et al. 1975
Condylarthra - Paroxyclaenidae	cf. Paroxyclaenidae indet.¹¹, cf. Paroxyclaenus sp.¹¹ cf. Paroxyclaenidae indet.¹¹ Weitzel placental cf. Paroxyclaenus sp.¹¹ Teilhard de Chardin 1922 placental MNHN Av 5714, 5909
Primates - Paromomyidae	Phenacolemur lapparenti n. sp.⁸, Arcius fuscus⁸ Phenacolemur lapparenti n. sp.⁸ Russell et al. 1967 primate MNHN AV 5849 type Arcius fuscus⁸ Russell et al. 1967 primate
Primates - Notharctidae	Cantius savagei⁵ Cantius savagei⁵ Gingerich 1977 primate
Primates - Plesiadapidae	Platychoerops daubrei⁴ Platychoerops daubrei⁴ Lemoine 1880 primate specimens are in the MNHN and/or the Louis private collection
Primates - Toliapinidae	Toliapina vinealis n. gen. n. sp.⁶, Avenius amatorum n. gen. n. sp.¹³ Toliapina vinealis n. gen. n. sp.⁶ Hooker et al. 1999 primate MNHN AV-700-Ph - holotype Avenius amatorum n. gen. n. sp.¹³ Russell et al. 1992 primate MNHN AV-1093-Ph - holotype
Creodonta -	Morlodon vellerei¹⁴ Morlodon vellerei¹⁴ Solé 2013 creodont MNHN Av 4785, 5917, (50)4711, 290, 5012, 467, 5070 and 5773
Creodonta - Hyaenodontidae	cf. Preregidens langebadrae¹⁶ cf. Preregidens langebadrae¹⁶ Solé et al. 2015 creodont MNHN.F.Av5773; Rich (1971) referred the specimen to ‘cf. Deltatheridia or Carnivora’
Theriamorpha -	Uintacyon hookeri¹⁵ Uintacyon hookeri¹⁵ Solé 2014 placental MNHN.F.Av5005, Av5006, Av5734, Av5817, Av4969, Av5820, Av4934, Av229BN, Av4756, Av15820
Carnivora - Miacidae	Miacidae indet.¹¹ Miacidae indet.¹¹ Cope 1880 carnivoran MNHN Av 5895
Theriamorpha - Viverravidae	cf. Viverravus sp.¹¹ cf. Viverravus sp.¹¹ Marsh 1872 placental MNHN Av 4916
Artiodactyla - Choeropotamidae	Cuisitherium sp.¹⁷ Cuisitherium sp.¹⁷ Sudre et al. 1983 even-toed ungulate
Artiodactyla - Diacodexeidae	Diacodexis varleti¹⁷, Bunophorus cappettai¹⁷ Diacodexis varleti¹⁷ Sudre 1983 even-toed ungulate Bunophorus cappettai¹⁷ even-toed ungulate
Theriamorpha - Nyctitheriidae	Placentidens lotus¹² Placentidens lotus¹² Russell et al. 1973 placental
Chiroptera - Icaronycteridae	Icaronycteris menui¹² Icaronycteris menui¹² Russell et al. 1973 bat
Chiroptera - Natalidae	cf. Ageina sp.¹² cf. Ageina sp.¹² Russell et al. 1973 funnel-eared bat
Chiroptera - Archaeonycteridae	Archaeonycteris brailloni n. sp.¹² Archaeonycteris brailloni n. sp.¹² Russell et al. 1973 bat type locality
Cimolesta - Sarcodontidae	cf. Hyracolestes sp.¹¹ cf. Hyracolestes sp.¹¹ Matthew and Granger 1925 eutherian MNHN Av 4614
Cimolesta - Apatemyidae	Apatemys mutiniacus n. sp., Apatemys sigogneaui n. sp., Heterohyus "sp. I", Heterohyus "sp. II", Heterohyus "sp. III" Apatemys mutiniacus n. sp. Russell et al. 1979 eutherian Apatemys sigogneaui n. sp. Russell et al. 1979 eutherian Heterohyus "sp. I" Gervais 1848 eutherian Heterohyus "sp. II" Gervais 1848 eutherian Heterohyus "sp. III" Gervais 1848 eutherian
Cimolesta - Esthonychidae	"Franchaius minimus" = Plesiesthonyx minimus⁵ "Franchaius minimus" = Plesiesthonyx minimus⁵ Baudry 1992 tillodont
Theriamorpha - Leptictidae	cf. Didelphodus sp.¹¹ cf. Didelphodus sp.¹¹ Cope 1882 eutherian MNHN Av 4917

¹Augé 1990; ²Augé 2005; ³Broin 1977; ⁴Gingerich 1976; ⁵Hooker 2010; ⁶Hooker et al. 1999; ⁷Lapparent de Broin 2003; ⁸López-Torres and Silcox 2018; ⁹Rage 1973; ¹⁰Rage 1977; ¹¹Rich 1971; ¹²Russell et al. 1973; ¹³Russell et al. 1992; ¹⁴Solé 2013; ¹⁵Solé 2014; ¹⁶Solé et al. 2015; ¹⁷Sudre and Erfurt 1996