Laño L1A (Cretaceous of Spain)

Also known as Laño Quarry

Where: Castilla y Leon, Spain (42.7° N, 2.6° W: paleocoordinates 33.0° N, 0.1° W)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Vitoria Formation, Late/Upper Campanian (83.5 - 70.6 Ma)

• in a sector of the south limb of the Miranda-Trevino syncline, level is also called L1A

• bed-level stratigraphic resolution

Environment/lithology: coastal; hardground, conglomeratic sandstone and black siltstone

• upper foreshore and backshore environments to a subcontinental, plant-rich perhaps swampy environment. The vertebrate bearing levels indicate a continental depositional environment affected by high-energy episodes, a fluviatile environment.
• Multidirectional cross-laminated sands with microconglomerates in the bottom-sets, rarely more than 1m thick. Strata become massive toward the top, and are frequently intersected by abrasion surfaces showing an iron-mineralized crust or hard-ground. The most important bone accumulation is between two of these hardgrounds. The unit is topped by two black silt beds.

Size classes: macrofossils, mesofossils, microfossils

Preservation: permineralized, replaced with pyrite

Collection methods: sieve,

• mostly chelonian shell fragments (30)

Primary reference: J. L. Sanz. 1986. Nouveaux gisements de dinosaures dans le Crétacé espagnol [New dinosaur localities in the Spanish Cretaceous]. Les Dinosaures de La Chine à La France, Colloque International de Paléontologie, Toulouse, France, 2-6 Septembre 1985; Muséum d'Histoire Naturelle de Toulouse, Toulouse 81-88 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 28406: authorized by John Alroy, entered by Erin Leckey on 12.02.2003, edited by Matthew Carrano and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

 Elopiformes - Phyllodontidae
Phyllodontidae indet.11 Sauvage 1875
 Amiiformes - Palaeolabridae
? Palaeolabrus sp.11 Casier 1967 bowfin
 Lepisosteiformes - Lepisosteidae
Atractosteus sp.11 Rafinesque 1820 gar
 Temnospondyli - Albanerpetontidae
Albanerpetontidae indet.5 Fox and Naylor 1982 tetrapod
 Salientia -
Anura indet.5 frog
distal part of 2 humeri, 8 ilia, fragmentary maxillaries, 2 broken sphenethmoids
 Salientia - Palaeobatrachidae
Palaeobatrachidae indet.5 Cope 1865 frog
43 prearticulars, 1 presacral vert, 2 synsacrums, 6 distal parts of humeri
 Salientia - Discoglossidae
Discoglossidae indet.1 Günther 1859 painted frog
6 prearticulars, 5 presacral verts, 7 sacral verts, 1 urostyle, 7 ilia, 2 humeri
 Caudata -
Caudata indet.5 salamander
may be Salamandra
 Testudines - Dortokidae
Dortoka vasconica4 Lapparent de Broin and Murelaga 1996 turtle
 Testudines - Bothremydidae
Polysternon atlanticum4 Lapparent de Broin and Murelaga 1996 sideneck turtle
? Elochelys sp.4 Nopsca 1931 sideneck turtle
right partial epiplastron
 Testudinata - Helochelydridae
Solemys vermiculata n. sp.4 Lapparent de Broin and Murelaga 1996 turtle
hypodigme around 260 shell fragments, most of them are peripherals and pleural fragments, around 25 fragments of plastron including the hologype
Solemys vermiculata9 Lapparent de Broin and Murelaga 1996 turtle
Helochelydra nopcsai4 Lapparent de Broin and Murelaga 1999 turtle
partial shell, lots of fragments
 Saurischia -
Lirainosaurus astibiae n. gen. n. sp.11
Lirainosaurus astibiae n. gen. n. sp.11 Sanz et al. 1999 sauropod
orig. 1 tooth
 Avetheropoda -
Coelurosauria indet.3 Huene 1914 coelurosaur
 Avetheropoda - Dromaeosauridae
Dromaeosauridae indet.11 Colbert and Russell 1969 coelurosaur
 Avetheropoda - Troodontidae
cf. Euronychodon sp.11 Antunes and Sigogneau-Russell 1991 coelurosaur
 Avetheropoda -
? Ornithomimosauria indet.3 Barsbold 1976 ornithomimosaur
 Saurischia -
Neoceratosauria indet.11 Novas 1991 ceratosaur
femora, MHNV
 Saurischia - Abelisauridae
? Abelisauridae indet.1 Bonaparte and Novas 1985 ceratosaur
 Ornithischia - Nodosauridae
Nodosauridae indet.1 Marsh 1890 ornithischian
cf. Struthiosaurus austriacus10 Bunzel 1871 ornithischian
a partial skeleton, including the synsacrum, portions of the pelvic girdle and hindlimbs, a dorsal vertebra, dorsal rib fragments and a dermal scute
 Neornithischia - Hadrosauridae
Hadrosauridae indet.11 Cope 1869 hadrosaurid
MCNA 10510, tooth
 Neornithischia - Rhabdodontidae
Rhabdodon sp.12 Matheron 1869 rhabdodontid
 Pterosauria - Azhdarchidae
cf. Azhdarcho sp.11 Nessov 1981 pterosaur
 Loricata - Allodaposuchidae
Allodaposuchus precedens2 Nopcsa 1928 crocodilian
may be Allodaposuchus
Musturzabalsuchus buffetauti11 Buscalioni et al. 1997 crocodilian
 Loricata - Hylaeochampsidae
Acynodon iberoccitanus11 Buscalioni et al. 1997 crocodilian
 Loricata -
Ischyrochampsa sp.11 Vasse 1995 crocodilian
 Squamata - Iguanidae
Iguanidae indet.13 Oppel 1811 squamates
 Squamata - Madtsoiidae
"Madtsoia laurasiae" = Menarana laurasiae13, Herensugea caristiorum13
"Madtsoia laurasiae" = Menarana laurasiae13 Rage 1996 snake
Herensugea caristiorum13 Rage 1996 snake
 Squamata -
Lacertilia "indet. 1"13 lizard
numerous remains (teeth)
Lacertilia "indet. 2"13 lizard
Lacertilia "indet. 3"13 lizard
Lacertilia indet.13 lizard
Scincomorpha indet.13 Camp 1923 lizard
Amphisbaenia indet.1 Gray 1844 worm lizard
? Amphisbaenia indet.13 Gray 1844 worm lizard
10 vertebrae
 Tribosphenida -
Eutheria indet.7 eutherian
 Tribosphenida - Zhelestidae
Lainodon " new species "8 Gheerbrant and Astibia 1994 eutherian
L1AT 18 (talonid of left M/3) and L1AT 13 and (talonid of right M/1 or M/2)
Lainodon orueetxebarriai n. gen. n. sp.6 Gheerbrant and Astibia 1994 eutherian
Lainodon cf. orueetxebarriai8 Gheerbrant and Astibia 1994 eutherian
L1AT 20 (distal part of right lower premolar)
Lainodon orueetxebarriai8 Gheerbrant and Astibia 1994 eutherian
new species
Lainodon ragei n. sp.8 Gheerbrant and Astibia 2012 eutherian
may be Marsupialia
 Theriiformes -
Theria "indet. B"1 Parker and Haswell 1897 mammal