Level 3, Benguérir (Cretaceous of Morocco)

Where: Morocco (32.3° N, 7.8° W: paleocoordinates 24.4° N, 8.4° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Late/Upper Maastrichtian (70.6 - 66.0 Ma)

• The use of dissimilarity indices and agglomerative method underscores two distinct associations: a lower one including the levels L6 and L5, and an upper one comprising the levels L4 to L2. These two associations allow to separate a lower and an upper Maastrichtian level. Moreover, the special morphology of the teeth of S. pristodontus occurring in L2, compared to what is observed in lower levels, permits us to distinguish a lower Upper Maastrichtian and an upper Upper Maastrichtian faunal assemblage.

• group of beds-level stratigraphic resolution

Environment/lithology: marine; poorly lithified phosphorite

Size classes: macrofossils, mesofossils

Reposited in the MNHN

Collection methods: surface (in situ),

• Repository: University of Montpellier 2; Office Chérifien des Phosphates, Khouribga, Morocco (OCP)

Primary reference: H. Cappetta, N. Bardet, X. Pereda Suberbiola, S. Adnet, D. Akkrim, M. Amalik, and A. Benabdallah. 2014. Marine vertebrate faunas from the Maastrichtian phosphates of Benguérir (Ganntour Basin, Morocco): Biostratigraphy, palaeobiogeography and palaeoecology. Palaeogeography, Palaeoclimatology, Palaeoecology 409:217-238 [M. Clapham/M. Clapham]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 213665: authorized by Matthew Clapham, entered by Matthew Clapham on 07.09.2020

Creative Commons license: CC BY (attribution)

Taxonomic list

Chondrichthyes
 Myliobatiformes - Rhombodontidae
Rhombodus meridionalis Arambourg 1952 stingray
Rhombodus microdon Arambourg 1952 stingray
 Myliobatiformes -
Ixobatis mucronata Arambourg 1952 stingray
Coupatezia elevata Noubhani and Cappetta 1997 stingray
Dasyrhombodus bondoni Arambourg 1952 stingray
 Rajiformes - Rhinobatidae
"Rhinobatos" "sp. 1" Link 1790 guitarfish
 Rajiformes - Sclerorhynchidae
Ctenopristis nougareti Arambourg 1940 ray
Ganopristis leptodon Arambourg 1935 ray
 Galeomorphi -
 Orectolobiformes - Ginglymostomatidae
Plicatoscyllium gharbii Noubhani and Cappetta 1997 nurse shark
Ginglymostoma sp. Müller and Henle 1838 nurse shark
 Lamniformes - Aquilolamnidae
Cretomanta sp. Case et al. 1990 mackerel shark
 Lamniformes - Serratolamnidae
Serratolamna serrata Agassiz 1838 mackerel shark
 Lamniformes - Anacoracidae
Squalicorax pristodontus Agassiz 1835 crow shark
Squalicorax microserratus2 Cappetta et al. 2014 crow shark
 Lamniformes - Otodontidae
Cretolamna maroccana mackerel shark
Reptilia
 Squamata - Mosasauridae
Halisaurus arambourgi Bardet et al. 2005 mosasaur
Prognathodon sp. Dollo 1889 mosasaur
Prognathodon currii1 Christiansen and Bonde 2002 mosasaur
OCP.DEK/GE (two teeth)
Globidens phosphaticus n. sp.1 Bardet et al. 2005 mosasaur
OCP.DEK/GE 337 - type (mid-posterior tooth); paratypes: OCP.DEK/GE 338 (an anterior tooth) and OCP.DEK/GE 339-343 (all median teeth); referred material: OCP.DEK/GE unnumbered (21 teeth)
Carinodens minalmamar n. sp.4 Schulp et al. 2009 mosasaur
OCP DEK/GE 453
Eremiasaurus heterodontus LeBlanc et al. 2012 mosasaur
"Platecarpus" ptychodon Arambourg 1952 mosasaur
 Squamata - Pachyvaranidae
Pachyvaranus crassispondylus3 Arambourg 1952 squamates
OCP.DEK/GE (vertebrae)
 Plesiosauria - Elasmosauridae
Elasmosauridae indet. Cope 1869 elasmosaur
 Testudines - Bothremydidae
Bothremydidae indet. sideneck turtle
 Testudines -
Chelonioidea indet. Baur 1893 sea turtle
Actinopteri
 Aulopiformes - Dercetidae
 Aulopiformes - Enchodontidae