Phosphorites du Quercy (for taxa with non-specific locality data) (Eocene to of France)

Also known as Quercy Phosphorites

Where: Languedoc-Roussillon-Midi-Pyrénées, France (44.3° N, 1.6° E: paleocoordinates 43.9° N, 1.1° W)

• coordinate estimated from map

• basin-level geographic resolution

When: Quercy Phosphorites Formation, Late/Upper Eocene to Late/Upper Eocene (37.2 - 16.0 Ma)

• General collection for specimens from the Phosphorites du Quercy with no specific locality or age data, particularly from older museum collections.

• formation-level stratigraphic resolution

Environment/lithology: terrestrial; phosphorite

Size classes: macrofossils, mesofossils

Primary reference: M. L. Augé. 2005. Evolution des lézards du Paléogène en Europe. Mémoires du Muséum national d'histoire naturelle 192:1-369 [P. Barrett/T. Cleary/T. Cleary]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 179194: authorized by Paul Barrett, entered by Terri Cleary on 03.06.2016, edited by Patricia Holroyd, Mark Uhen and Grace Varnham

Creative Commons license: CC BY (attribution)

Taxonomic list

Aves
 Cariamiformes - Cariamidae
Ameghinornis minor n. gen.2 Gaillard 1939 seriema
 Strigiformes - Sophiornithidae
Sophiornis quercynus n. gen. n. sp.3, Palaeobyas cracrafti n. gen. n. sp.3, Palaeotyto cadurcencis n. gen. n. sp.3
Sophiornis quercynus n. gen. n. sp.3 Mourer-Chauviré 1987 owl
Palaeobyas cracrafti n. gen. n. sp.3 Mourer-Chauviré 1987 owl
Palaeotyto cadurcencis n. gen. n. sp.3 owl
 Strigiformes - Strigidae
"Selenornis henrici n. gen." = Otus henrici3, "Necrobyas medius n. sp." = Prosybris medius3
"Selenornis henrici n. gen." = Otus henrici3 Milne-Edwards 1892 true owl
"Necrobyas medius n. sp." = Prosybris medius3 Mourer-Chauviré 1987 true owl
Reptilia
 Crocodylia - Alligatoridae
Alligator gaudryi n. sp.5 de Stefano 1905 alligator
 Testudines - Testudinidae
Testudo phosphoritarum n. sp.5 Rage 1988 turtle
 Squamata -
Squamata indet.1 squamates
A ?sacral vertebra (NHMW 2019/0095/0002).
Anguimorpha indet.1 squamates
Five presacral vertebrae (NHMW 2019/0046/0003- NHMW 2019/0046/0007); a partial pectoral girdle (NHMW 2019/0095/0001).
 Squamata - Palaeovaranidae
Palaeovaranus sp.1 Zittel 1887 squamates
A left maxilla (NHMW 2019/0048/0002); a left dentary (NHMW 2019/0058/0054); two right dentaries (NHMW2019/0058/0055 and NHMW 2019/0058/0056); 35 presacral verte- brae (NHMW 2019/0058/0001-NHMW 2019/0058/0009, NHMW 2019/0058/0011-NHMW 2019/0058/0027, NHMW 2019/0058/0030, NHMW 2019/0058/0041-NHMW 2019/0058/0046, NHMW 2019/0058/0051, and NHMW 2019/0067/0001); two articulated sacral vertebrae (NHMW 2019/0058/0028); two sacral vertebrae (NHMW 2019/0058/0029 and NHMW 2019/0058/0047); 16 caudal vertebrae (NHMW 2019/0058/0031-NHMW 2019/0058/0040, NHMW 2019/0058/0048-NHMW 2019/0058/0050, NHMW 2019/0058/0052, NHMW 2019/0058/0053, and NHMW 2019/0067/0002).
Palaeovaranus lismonimenos n. sp.1 Georgalis et al. 2021 squamates
Holotype: An almost complete parietal (NHMW 2019/0047/0001); A complete parietal (MNHN.F.QU17177) and a partial parietal (UM BFI 1873), both from juvenile individuals. Tentatively also, two frontals (MNHN.F.QU17175 and UM PRA 8).
Palaeovaranus cayluxi1 Zittel 1887 squamates
Two almost complete parietals (NHMW 2019/0048/0001 and MNHN.F.QU17176).
 Squamata -
Saniwa sp.1 Leidy 1870 squamates
Three presacral vertebrae (NHMW 2019/0065/0001-NHMW 2019/0065/0003).
 Squamata - Anguidae
Anguinae indet.1 Gray 1825 squamates
Two presacral vertebrae (NHMW 2019/0093/0001 and NHMW 2019/0093/0002).
Placosaurus sp. Gervais 1852 squamates
Part of the anterior of a right dentary, part of the anterior of a left dentary, a premaxilla. Likely mid-late Eocene in age. three presacral vertebrae (NHMW 2019/0046/0001, NHMW 2019/0046/0002, and possibly also NHMW 2019/0046/0008); one caudal vertebra (NHMW 2019/0046/0009)
Placosaurus rugosus Gervais 1852 squamates
MNHN QU 16564, complete right mandible (holotype of Placosaurus europaeus in Filhol 1877); some osteoderms(?); short maxilla fragment; MNHN QU 17178, complete right mandible. Other collections are mid-late Eocene in age.
Melanosaurini indet.1 Sullivan 1979 squamates
Four presacral vertebrae (NHMW 2019/0094/0001- NHMW 2019/0094/0003 and NHMW 2019/0094/0005); one sacral vertebra (NHMW 2019/0094/0004).
 Squamata - Necrosauridae
Necrosaurus eucarinatus Kuhn 1940 squamates
MNHN QU 17177, one frontal, two parietals. Likely Eocene age.
Necrosaurus cayluxi n. gen. n. sp. Filhol 1873 squamates
MNHN fig 434 in Filhol 1877 (holotype), dentary; MNHN QU 17172, right dentary; three incomplete right dentaries; one incomplete left dentary; two incomplete maxillae (one left, one right); fragment of jaw; one premaxilla; MNHN QU 17176, parietal; two additional parietals; MNHN QU 17175, frontal; two additional frontals; QU 17173, part of the posterior of a mandible; QU 17174, right quadrate; a number of dorsal and caudal vertebrae (BMNH). Some material from BSPHM collections in Fejervary 1935. Likely late Eocene in age.
 Squamata -
Eurheloderma gallicum n. gen. n. sp.
Eurheloderma gallicum n. gen. n. sp. Hoffstetter 1957 squamates
MNHN QU 17746 (holotype), right maxilla; additional MNHN material: five maxillae, one left dentary, one left pterygoid, two parietals, five dorsal vertebrae, two posterior cervical vertebrae, two lumbar vertebrae; BMNH R3487, a right dentary; EMP collections: one parietal; FSM coll.: one parietal. Late Eocene (or possibly Oligocene?).
 Squamata - Cordylidae
Pseudolacerta quercyini Augé 2005 spinytail lizard
?a fragmentary left dentary, ?a fragmentary right dentary. Probably mid-late Eocene in age.
Pseudolacerta mucronata n. gen. n. sp. Filhol 1877 spinytail lizard
Holotype (MNHN, presumed lost), one dentary; MNHN QU 17159, right dentary with broken posterior; additional three right dentaries. Most likely from the late Eocene.
 Squamata - Scincidae
Ayalasaurus tenuis n. gen. n. sp.
Ayalasaurus tenuis n. gen. n. sp. Augé 2005 skink
MNHN QU 17190 (holotype), left dentary with splenial and start of angular; left dentary; MNHN QU 17191,three right dentaries. Certainly from the late Eocene.
 Squamata - Gekkonidae
Cadurcogekko piveteaui n. gen. n. sp. Hoffstetter 1946 gecko
MNHN QU 17163 and QU 17164 (syntypes), left dentary and incomplete maxilla, respectively; seven dentaries, two maxillae, one frontal. Probably mid-late Eocene in age.
Cadurcogekko cf. piveteaui1 Hoffstetter 1946 gecko
A frontal (NHMW 2019/0052/0002); a left maxilla (NHMW 2019/0052/0001).
 Squamata -
"Placosaurus quercyi n. sp." = Paraplacosauriops quercyi, "Paraplacosauriops quercyi" = Paraplacosauriops quercyi1
"Placosaurus quercyi n. sp." = Paraplacosauriops quercyi Filhol 1882 squamates
MNHN QU 16569 (neotype), right dentary; QU 16565, two left dentaries; QU 17730, three right dentaries; a mandible; two incomplete maxillae. Certainly from the middle or late Eocene. Additional: YPM-PU 11494 from reference 59390, nearly complete right maxilla: it is said to come from "Caylus, Tarn-et-Garonne", but this could mean it is from a number of established Quercy localities.
"Paraplacosauriops quercyi" = Paraplacosauriops quercyi1 Filhol 1882 squamates
An incomplete left maxilla (NHMW 2019/0049/0003); a right dentary (NHMW 2019/0049/0001); a left dentary (NHMW 2019/0049/0002).
 Squamata - Colubridae
Natrix mlynarskii n. sp.4 Rage 1988 water snake
MNHN QU 17181 (holotype), one mid-trunk vertebra; MNHN QU 17182, one mid-trunk vertebra; Rage notes that since a Mas de Got specimen exists these specimens are most likely Oligocene in age
 Squamata - Boidae
Palaeopython cadurcensis n. sp.5 Filhol 1877 boa
Palaeopython filholi n. gen. n. sp.5 de Rochebrune 1880 boa
MNHN QU 322 (holotype), vertebra (see also Rage 1974, ref. 16854)
Palaeopython neglectus n. sp.5 de Rochebrune 1884 boa
Paleryx cayluxi n. sp.5 De Stefano 1905 boa
Plesiotortrix edwardsi n. sp.5 de Rochebrune 1884 boa
 Squamata -
Uromastyx europaeus n. sp. Moody 1980 spiny-tailed lizard
MNHN No. QU 17160 (holotype), right dentary; MNHN QU 17161, right and left dentaries; MNHN QU 17183-17189, three incomplete dentaries and four incomplete maxillae. Most likely from the Oligocene.
 Squamata - Agamidae
Quercygama galliae n. gen. n. sp.
Quercygama galliae n. gen. n. sp. Filhol 1877 squamates
MNHN QU16560 (holotype), mandible. Most likely from the early Oligocene.
 Squamata - Iguanidae
Geiseltaliellus lamandini n. sp. Filhol 1877 squamates
MNHN No. QU 17739 (holotype), incomplete right mandible with dentary, splenial, coronoid and small part of supra-angular and angular; additional material: seven right dentaries, three left dentaries, and ?a frontal (MNHN QU 17162). Most likely late Eocene in age.
 Squamata - Amphisbaenidae
Omoiotyphlops priscus n. gen. n. sp.
Omoiotyphlops priscus n. gen. n. sp. de Rochebrune 1884 worm lizard
MNHN QU 16573 (holotype), five associated vertebrae.
 Squamata - Lacertidae
Mediolacerta sp.1 Augé 2005 squamates
A left dentary (NHMW 2019/0050/0001).
Mediolacerta roceki Augé 2005 squamates
twelve right dentaries, five left dentaries, a left dentary with splenial; MNHN QU 17728, a mostly complete right mandible. Most likely Oligocene in age.
Pseudeumeces sp.1 Hoffstetter 1944 squamates
A left maxilla (NHMW 2019/0051/0004); a left dentary (NHMW 2019/0051/0003); a right dentary (NHMW 2019/0051/0005).
Pseudeumeces cadurcensis n. gen. n. sp. Filhol 1877 squamates
MNHN possibly lost holotype (figured in Filhol 1877), dentary; MNHN QU 17169, complete right mandible; left dentary; six maxillae; MNHN QU 17180, hundred dentaries. Probably Oligocene in age.
Pseudeumeces kyrillomethodicus n. sp.1 Georgalis et al. 2021 squamates
Holotype: NHMW 2019/0051/0001, left dentary; NHMW 2019/0051/0002, right dentary
Plesiolacerta lydekkeri Hoffstetter 1942 squamates
MNHN QU 17179, seven left dentaries; MNHN QU 17727, four right dentaries; QU 17166, sixteen left maxillae; [no cat no.] twelve right maxillae; QU 17167, eleven dorsal vertebrae; two cervical vertebrae, one caudal vertebra; QU 17168, three parietals, QU 17745, one premaxilla. Age range of other collections: mid Eocene-early Oligocene.
Quercycerta maxima Augé 2005 squamates
MNHN QU 17729 (paratype), right dentary; additional material: three left dentaries, five right dentaries, fragmentary left maxilla. Other collections are mid-Eocene in age.
Dracaenosaurus croizeti Gervais 1848 squamates
MNHN 1903-20, left dentary (previously holotype for Dracaenosaurus sauvegi). Other collections are from the late Oligocene.
 Squamata - Teiidae
Brevisaurus smithi n. gen. n. sp.
Brevisaurus smithi n. gen. n. sp. Augé 2005 squamates
MNHN QU 17742 (holotype), right dentary; QU 17741, right maxilla; additional fragment of right maxilla, left dentary, partial right dentary. Other collections are mid-Eocene in age.
Amphibia
 Caudata - Urodela
Megalotriton filholi n. sp.5 Zittel 1890 newt
Megalotriton portisi n. sp.5 De Stefano 1903 newt
Phosphotriton sigei n. gen. n. sp.6 Tissier et al. 2016 newt
MNHN.F.QU 17755 (holotype)
 Caudata -
Heteroclitotriton zitteli n. sp.5 De Stefano 1903 salamander
 Salientia - Bufonidae
Bufo serratus n. sp.5
Bufo serratus n. sp.5 Filhol 1876 toad
 Salientia -
"Rana plicata n. sp." = Thaumastosaurus gezei5, Thaumastosaurus bottii n. sp.5
"Rana plicata n. sp." = Thaumastosaurus gezei5 Rage and Rocek 2007 frog
Thaumastosaurus bottii n. sp.5 De Stefano 1903 frog