PBDB Collection

Lull 2 Quarry (UCMP V-5620) (Cretaceous of the United States)

Also known as Lull's Locality 2

Where: Niobrara County, Wyoming (43.1° N, 104.6° W: paleocoordinates 49.6° N, 79.5° W)

• coordinate estimated from map

• small collection-level geographic resolution

When: Lance Formation, Late/Upper Maastrichtian (70.6 - 66.0 Ma)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; carbonaceous sandstone

• "sandstone + lignites"

Size class: mesofossils

Collected by W. A. Clemens; reposited in the UCMP

Collection methods: bulk, surface (float), sieve,

Primary reference: W. A. Clemens. 1964. Fossil mammals of the type Lance Formation, Wyoming. Part I. Introduction and Marsupialia. University of California Publications in Geological Sciences 48:1-105 [J. Alroy/J. Alroy/J. Alroy]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 14585: authorized by John Alroy, entered by John Alroy on 26.07.1994, edited by Matthew Carrano, Roger Benson and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

• this is NOT Hatcher's locality No. 2, which is UCMP V-5815

Show authors, comments, and common names

Gastropoda
Heterostropha - Physidae	cf. Physa sp.⁹ cf. Physa sp.⁹ Draparnaud 1801 snail
Architaenioglossa - Viviparidae	cf. Lioplacodes sp.⁹, cf. Viviparus sp.⁹ cf. Lioplacodes sp.⁹ Meek and Hayden 1864 snail cf. Viviparus sp.⁹ Montfort 1810 snail
Sorbeoconcha - Hydrobiidae	Reesidella sp.⁹ Reesidella sp.⁹ Yen 1952 snail
Bivalvia
Unionida - Unionidae	Unionidae indet.⁹ Unionidae indet.⁹ Rafinesque 1820 freshwater mussel
Cardiida - Corbiculidae	? Corbiculidae indet.⁹ ? Corbiculidae indet.⁹ Gray 1847 clam
Mammalia
Theriamorpha - Pediomyidae	"Pediomys florencae" = Protolambda florencae⁴, "Pediomys hatcheri" = Protolambda hatcheri⁴, "Pediomys krejcii" = Leptalestes krejcii⁴, "Pediomys cooki" = Leptalestes cooki⁴, Pediomys elegans⁴ "Pediomys florencae" = Protolambda florencae⁴ Clemens 1966 metatherian "Pediomys hatcheri" = Protolambda hatcheri⁴ Osborn 1898 metatherian "Pediomys krejcii" = Leptalestes krejcii⁴ Clemens 1966 metatherian this IS the type locality according to Clemens 1973 "Pediomys cooki" = Leptalestes cooki⁴ Clemens 1966 metatherian Pediomys elegans⁴ Marsh 1889 metatherian
Theriamorpha - Herpetotheriidae	"Nortedelphys intermedius" = Nortedelphys jasoni³, Nortedelphys minimus³ "Nortedelphys intermedius" = Nortedelphys jasoni³ Storer 1991 marsupial Nortedelphys minimus³ Case et al. 2004 marsupial
Theriamorpha - Stagodontidae	Didelphodon vorax⁴ Didelphodon vorax⁴ Marsh 1889 metatherian
Polydolopimorphia - Glasbiidae	Glasbius intricatus⁴ Glasbius intricatus⁴ Clemens 1966 metatherian
Theriamorpha - Alphadontidae	Alphadon marshi⁴, "Alphadon rhaister" = Turgidodon rhaister Alphadon marshi⁴ Simpson 1927 metatherian includes A. marshi sensu stricto according to Clemens 1973; A. "wilsoni" present according to Lillegraven 1969 "Alphadon rhaister" = Turgidodon rhaister Clemens 1966 metatherian
Theriamorpha -	"Alphadon lulli n. sp." = Protalphadon lulli "Alphadon lulli n. sp." = Protalphadon lulli Clemens 1966 metatherian
Deltatheroida -	cf. Deltatheroides sp.¹⁴ cf. Deltatheroides sp.¹⁴ Gregory and Simpson 1926 metatherian two unidentified therian teeth figured by Clemens 1973
Theriamorpha - Gypsonictopidae	Gypsonictops hypoconus⁵ Gypsonictops hypoconus⁵ Simpson 1927 eutherian
Cimolesta - Cimolestidae	Cimolestes incisus⁵, Cimolestes stirtoni⁵ Cimolestes incisus⁵ Marsh 1889 eutherian Cimolestes stirtoni⁵ Clemens 1973 eutherian
Multituberculata - Eucosmodontidae	Clemensodon megaloba n. gen. n. sp.²¹ Clemensodon megaloba n. gen. n. sp.²¹ Krause 1992 multituberculate
Multituberculata -	Cimexomys minor⁵ Cimexomys minor⁵ Sloan and Van Valen 1965 multituberculate six M1 specimens of "Mesodma sp." of Clemens 1964a
Multituberculata - Neoplagiaulacidae	Mesodma hensleighi²⁶, Mesodma formosa, Mesodma thompsoni n. sp. Mesodma hensleighi²⁶ Lillegraven 1969 multituberculate Mesodma formosa Marsh 1889 multituberculate confirmed by Novacek and Clemens 1977 Mesodma thompsoni n. sp. Clemens 1964 multituberculate
Multituberculata - Cimolodontidae	Cimolodon nitidus Cimolodon nitidus Marsh 1889 multituberculate
Multituberculata - Cimolomyidae	Meniscoessus robustus, Cimolomys gracilis Meniscoessus robustus Marsh 1889 multituberculate Cimolomys gracilis Marsh 1889 multituberculate
Reptilia
Squamata -	"Sauriscus cooki n. gen. n. sp." = Estescincosaurus cooki⁹ "Sauriscus cooki n. gen. n. sp." = Estescincosaurus cooki⁹ Estes 1964 squamates UCMP 46088, 47811-47813, 47827-47828, 47831, 49994-49995, 50017-50019
Squamata - Chamopsiidae	Leptochamops denticulatus⁹, Chamops segnis⁹, Meniscognathus altmani n. gen. n. sp.⁹, Haptosphenus placodon n. gen. n. sp.⁹ Leptochamops denticulatus⁹ Gilmore 1928 squamates UCMP 46069, 47784, 47790, 47796-47798, 47800, 47825, 47803, 47806, 49901, 49904, 49906-49907, 49909, 49912, 49914, 49917 Chamops segnis⁹ Marsh 1892 squamates UCMP 46032-46033, 46094, 46100, 46614, 47782, 49833, 49871-49872, 49874, 49877-49878, 49880-49881, 49888, 50029, 54217 Meniscognathus altmani n. gen. n. sp.⁹ Estes 1964 squamates UCMP 46064, 46079, 46081, 46085, 46101, 46106, 46609-46610, 47834, 47846, 47848, 47853, 47859, 47865-47866 Haptosphenus placodon n. gen. n. sp.⁹ Estes 1964 squamates UCMP 46035, 49946-49947, 49949-49950, 50031
Squamata -	Coniophis precedens²³ Coniophis precedens²³ Marsh 1892 snake UCMP 45940, 46060, 46070, 50001‐50014, 53924‐ 53929, 54219 (vertebrae); UCMP 53935, 49998, 49999 (maxilla) and UCMP 49997, 50000 (dentaries)
Squamata - Iguanidae	Iguanidae indet.⁶ Iguanidae indet.⁶ Oppel 1811 squamates
Squamata - Xenosauridae	Exostinus lancensis⁹ Exostinus lancensis⁹ Gilmore 1928 squamates UCMP 46053, 46076, 46090, 46606, 47772, 47829, 50015-50016, 50024, 56091-56092
Squamata -	Paraderma bogerti⁹ Paraderma bogerti⁹ Estes 1964 squamates UCMP 49895, 49936, 49939, 49940
Squamata - Anguidae	"Peltosaurus piger" = Odaxosaurus piger⁹, cf. Gerrhonotus sp.⁹ "Peltosaurus piger" = Odaxosaurus piger⁹ Gilmore 1928 squamates UCMP 45935, 46017, 46072, 46089, 46096, 46607, 46611, 46624, 47753-47768, 49921-49933, 49959, 59992, 50030, 56056 cf. Gerrhonotus sp.⁹ Wiegmann 1828 alligator lizard UCMP 45938, 47766, 49931-49932, 49934
Squamata -	Parasaniwa wyomingensis⁹, Litakis gilmorei n. gen. n. sp.⁹, Colpodontosaurus cracens n. gen. n. sp.⁹, Cemeterius monstrosus²⁴, Varanoidea indet.¹⁶ Parasaniwa wyomingensis⁹ Gilmore 1928 squamates UCMP 46041, 46603, 46036, 47742, 47746, 47752, 49891-49892, 49900, 49935 Litakis gilmorei n. gen. n. sp.⁹ Estes 1964 squamates UCMP 47815 Colpodontosaurus cracens n. gen. n. sp.⁹ Estes 1964 squamates UCMP 46608, 49938, 49957 Cemeterius monstrosus²⁴ Longrich et al. 2012 squamates UCMP 49954, isolated tooth Varanoidea indet.¹⁶ Gray 1827 squamates UCMP 46051, 49952-49956
Testudinata -	Compsemys sp.¹⁹, Compsemys victa⁹ Compsemys sp.¹⁹ Leidy 1856 turtle Voucher: UCMP 72897 Compsemys victa⁹ Leidy 1856 turtle
Testudinata - Baenidae	? Baenidae indet.⁹, Baenidae sp.¹⁹, "Peckemys brinkman" = Cedrobaena brinkman²⁵, Plesiobaena sp.¹⁹, Plesiobaena antiqua¹⁵ ? Baenidae indet.⁹ Cope 1882 turtle Baenidae sp.¹⁹ Cope 1882 turtle Voucher: UCMP 72936 "Peckemys brinkman" = Cedrobaena brinkman²⁵ Lyson and Joyce 2009 turtle UCMP 49795 (skull) Plesiobaena sp.¹⁹ Gaffney 1972 turtle Voucher: UCMP 49759 Plesiobaena antiqua¹⁵ Lambe 1902 turtle UCMP 49759
Testudines - Pantrionychidae	"Plastomeninae sp." = Plastomenidae¹⁹, Helopanoplia sp.¹⁹ "Plastomeninae sp." = Plastomenidae¹⁹ Hay 1902 turtle Voucher: UCMP 173441 Helopanoplia sp.¹⁹ Hay 1908 turtle Voucher: UCMP 173440
Testudines - Chelydridae	cf. Chelydridae indet.⁶, Chelydridae sp.¹⁹ cf. Chelydridae indet.⁶ Swainson 1839 turtle Chelydridae sp.¹⁹ Swainson 1839 turtle Vouchers: UCMP 125776–788
Testudines - Trionychidae	Trionychidae sp.¹⁹, "Aspideretes beecheri" = Trionyx beecheri⁹ Trionychidae sp.¹⁹ Gray 1825 softshell turtle Vouchesr: UCMP 73028–29 "Aspideretes beecheri" = Trionyx beecheri⁹ Hay 1904 softshell turtle
Testudines - Emydidae	Emydinae indet.⁹ Emydinae indet.⁹ Rafinesque 1815 turtle "near Pseudemys?"
Testudines - Adocidae	Adocus sp.¹⁹ Adocus sp.¹⁹ Cope 1868 turtle Voucher: UCMP 173439
Testudines - Kinosternoidae	Kinosternoidae sp.¹⁹ Kinosternoidae sp.¹⁹ Holroyd and Hutchison 2002 turtle Voucher: UCMP 125762
Testudines - Nanhsiungchelyidae	Basilemys sp.⁹, Basilemys sp.¹⁹ Basilemys sp.⁹ Hay 1902 turtle Basilemys sp.¹⁹ Hay 1902 turtle Voucher: UCMP 72901
Loricata -	"Leidyosuchus sternbergi" = Borealosuchus sternbergii¹² "Leidyosuchus sternbergi" = Borealosuchus sternbergii¹² Gilmore 1910 crocodilian
Crocodylia - Alligatoridae	Brachychampsa montana¹² Brachychampsa montana¹² Gilmore 1911 crocodilian
Ornithischia - Ceratopsidae	cf. Triceratops sp.⁹ cf. Triceratops sp.⁹ Marsh 1889 ceratopsid
Ornithischia -	"Pachycephalosaurus grangeri" = Pachycephalosaurus wyomingensis⁶, Thescelosaurus sp.² "Pachycephalosaurus grangeri" = Pachycephalosaurus wyomingensis⁶ Gilmore 1931 ornithischian Thescelosaurus sp.² Gilmore 1913 ornithopod UCMP 124972-124977
Ornithischia - Hadrosauridae	"cf. Anatosaurus sp." = Edmontosaurus⁹ "cf. Anatosaurus sp." = Edmontosaurus⁹ Lambe 1917 hadrosaurine
Ornithischia - Ankylosauridae	Ankylosaurus magniventris² Ankylosaurus magniventris² Brown 1908 ankylosaurid UCMP 124399
Theropoda -	Theropoda indet.², ? Theropoda indet.⁹, Paronychodon lacustris⁹, cf. Paronychodon lacustris²⁷ Theropoda indet.² theropod UCM 45063 ? Theropoda indet.⁹ theropod AMNH 8113 Paronychodon lacustris⁹ Cope 1876 maniraptoran cf. Paronychodon lacustris²⁷ Cope 1876 maniraptoran UCMP 53283, 124400, 187086-7, 187089, 187091-3, 187096, 187098-100, 187102, 187108, 187112, 187114, 187120-1, 187123, 187126, 187128, 187130, 187133-4, 187156, 187162, 187164
Theropoda - Troodontidae	cf. Saurornithoides sp.⁹, Troodon sp.²⁷, Pectinodon bakkeri² cf. Saurornithoides sp.⁹ Osborn 1924 maniraptoran AMNH 8114 Troodon sp.²⁷ Leidy 1856 maniraptoran Pectinodon bakkeri² Carpenter 1982 maniraptoran UCMP 125240-125247, 187059, 187063, 187065-9, 187073-6
Theropoda - Dromaeosauridae	Richardoestesia cf. isosceles²⁷, cf. Zapsalis sp.²², Saurornitholestinae indet.²² Richardoestesia cf. isosceles²⁷ Sankey 2001 maniraptoran UCMP 213984, 187102 cf. Zapsalis sp.²² Cope 1876 maniraptoran UCM 45505; UCMP 187124-5, 187129, 187105 Saurornitholestinae indet.²² Longrich and Currie 2009 maniraptoran UCMP 187070, 124401, 187079
Aves
Charadriiformes - Cimolopterygidae	Ceramornis major n. gen. n. sp.¹, aff. Cimolopteryx sp.⁹, Cimolopteryx rara⁹ Ceramornis major n. gen. n. sp.¹ Brodkorb 1963 shorebird UCMP 53959 aff. Cimolopteryx sp.⁹ Marsh 1889 shorebird Cimolopteryx rara⁹ Marsh 1889 shorebird UCMP 53962, 53963, 53964
Hesperornithiformes -	Potamornis skutchi n. gen. n. sp.⁷ Potamornis skutchi n. gen. n. sp.⁷ Elzanowski et al. 2001 bird ? Hesperornithiformes, family unknown
Neornithes -	Neornithes "indet. sp. 3"²⁰, Gaviiformes indet.⁹ Neornithes "indet. sp. 3"²⁰ bird UCMP 53960 Gaviiformes indet.⁹ loon
Ciconiiformes -	Ciconiiformes indet.⁹ Ciconiiformes indet.⁹ Bonaparte 1854 bird
Pelecaniformes -	Torotix clemensi n. gen. n. sp.¹ Torotix clemensi n. gen. n. sp.¹ Brodkorb 1963 bird UCMP 53958
Neornithes -	Lonchodytes estesi n. gen. n. sp.¹, "Galloanseres indet." = Galloanserae⁸, Lonchodytes pterygius n. sp.¹ Lonchodytes estesi n. gen. n. sp.¹ Brodkorb 1963 bird UCMP 53954 "Galloanseres indet." = Galloanserae⁸ Sibley and Ahlquist 1990 bird UCMP 53969 Lonchodytes pterygius n. sp.¹ Brodkorb 1963 bird UCMP 53961
Reptilia
Theropoda -	cf. Dryptosaurus sp.⁹ cf. Dryptosaurus sp.⁹ Marsh 1877 coelurosaur
Theropoda - Tyrannosauridae	cf. Tyrannosaurus rex⁹, Aublysodon mirandus² cf. Tyrannosaurus rex⁹ Osborn 1905 tyrant lizard king Aublysodon mirandus² Leidy 1868 tyrannosaurid UCMP 124399
Theropoda - Ornithomimidae	cf. Ornithomimus sp.⁹ cf. Ornithomimus sp.⁹ Marsh 1890 ornithomimid
Pterosauria -	? Pterosauria indet.⁹ ? Pterosauria indet.⁹ pterosaur
Choristodera -	Champsosaurus sp.¹² Champsosaurus sp.¹² Cope 1876 choristodere
Amphibia
Temnospondyli - Albanerpetontidae	Albanerpetontidae indet.¹⁷, Albanerpeton nexuosus n. sp.¹¹ Albanerpetontidae indet.¹⁷ Fox and Naylor 1982 tetrapod UCMP 49534, 49536 Albanerpeton nexuosus n. sp.¹¹ Estes 1981 tetrapod UCMP 49547, 49535, 49538, 49540
Salientia - Pelobatidae	cf. Eopelobates sp.¹⁸ cf. Eopelobates sp.¹⁸ Parker 1929 spadefoot toad UCMP 55708-55709, 55711-55713 + 21 unnumbered specimens
Salientia -	Neobatrachia indet.⁹ Neobatrachia indet.⁹ frog 2 urostyles
Salientia - Hylidae	aff. Hylidae indet.⁹ aff. Hylidae indet.⁹ Rafinesque 1815 tree frog UCMP 55707, 55710; Neobatrachidae "near Hylidae?"
Salientia - Leptodactylidae	aff. Leptodactylidae indet.⁹ aff. Leptodactylidae indet.⁹ Stirton 1953 frog AMNH 8102, Neobatrachidae "near Leptodactylidae?"
Salientia -	Paradiscoglossus americanus¹³ Paradiscoglossus americanus¹³ Estes and Sanchíz 1982 frog UCMP 125830
Salientia - Palaeobatrachidae	Palaeobatrachus occidentalis n. sp.¹³ Palaeobatrachus occidentalis n. sp.¹³ Estes and Sanchíz 1982 frog UCMP 55704
Salientia -	Scotiophryne pustulosa¹⁰ Scotiophryne pustulosa¹⁰ Estes 1969 frog UCMP 55703
Proteida - Batrachosauroididae	Opisthotriton kayi⁹, "Prodesmodon copei n. gen. n. sp." = Prodesmodon copei⁹, "Cuttysarkus mcnallyi n. gen. n. sp." = Prodesmodon copei⁹ Opisthotriton kayi⁹ Auffenberg 1961 salamander UCMP 46063, 46078, 47817, 49501-49504, 49512-49514, 49520-49521, 49526, 49528-49532, 50021 "Prodesmodon copei n. gen. n. sp." = Prodesmodon copei⁹ Estes 1964 salamander UCMP 49509, 49511, 49533-49536, 49538-49540, 55775, 55777-55780, 55782-55783 "Cuttysarkus mcnallyi n. gen. n. sp." = Prodesmodon copei⁹ Estes 1964 salamander UCMP 46061, 46066, 46095, 49526-49527, 49941-49944, 49995, 54218
Caudata - Urodela	Scapherpeton tectum⁹, Lisserpeton bairdi¹², Habrosaurus dilatus⁹, Plethodontidae indet.⁶ Scapherpeton tectum⁹ Cope 1876 salamander UCMP 46507-49508, 49516-49517, 49541-49543, 49548, + >500 unnumbered elements Lisserpeton bairdi¹² Estes 1965 salamander Habrosaurus dilatus⁹ Gilmore 1928 siren UCMP 46013, 46043, 46046, 46054, 46062, 46065, 46073, 46077, 46083, 46097, 46104, 46107, 46460, 46612, 46613, 46627, 49491-49500, 49505-49506, 49518-49519, 49537, 55714-55717 Plethodontidae indet.⁶ Gray 1850 lungless salamander
Actinopteri
Amiiformes - Amiidae	"Kindleia fragosa" = Cyclurus fragosus⁹, Melvius cf. thomasi⁶, cf. Protamia sp.⁹ "Kindleia fragosa" = Cyclurus fragosus⁹ Jordan 1927 bowfin UCMP 46084, 47070-47072, 47075-47081, 54013-54014, 54025-54030, 54043-54044, 54047-54051, 54054-54056, 54059-54060, 54063-54068, 54083-54087, 54089-54091, 54095-54103, 54106-54120, 54141, 54144-54157, 54163-54166, 54174-54177, 54189, 54191-54195; + >105 unnumbered specimens Melvius cf. thomasi⁶ Bryant 1987 bowfin cf. Protamia sp.⁹ Leidy 1873 bowfin UCMP 56277; also listed as Amia sp.
Lepisosteiformes - Lepisosteidae	Lepisosteus occidentalis⁹ Lepisosteus occidentalis⁹ Leidy 1856 gar
Albuliformes - Albulidae	Coriops amnicolus¹² Coriops amnicolus¹² Estes 1969 bonefish UCMP 54167, 54169 + 1 unnumbered specimen
Elopiformes - Elopidae	Elopidae indet.⁹ Elopidae indet.⁹ Bonaparte 1846 scale fragments
Acanthomorphata - Sciaenidae	Platacodon nanus⁹ Platacodon nanus⁹ Marsh 1889 UCMP 46109, 53689-53709, 53711-53717; dentaries only (UCMPM 53708-53709, 53711-53711, 53717) referred to Estesesox foxi by Wilson et al. 1992
Aspidorhynchiformes - Aspidorhynchidae	Belonostomus longirostris⁹ Belonostomus longirostris⁹ Lambe 1902 UCMP 53979-53989
Osteichthyes
Acipenseriformes - Acipenseridae	Acipenser eruciferus⁹ Acipenser eruciferus⁹ Cope 1876 sturgeon UCMP 53904
Acipenseriformes - Polyodontidae	Paleopsephurus wilsoni¹² Paleopsephurus wilsoni¹² MacAlpin 1947 paddlefish
Chondrichthyes
Rajiformes - Sclerorhynchidae	Ischyrhiza avonicola n. sp.⁹ Ischyrhiza avonicola n. sp.⁹ Estes 1964 ray UCMP 53911, 53912, 53914, 53915, 53916, 53918, 62828
Rajiformes - Rhinobatidae	Myledaphus bipartitus⁹ Myledaphus bipartitus⁹ Cope 1876 guitarfish AMNH 9203-9213
Orectolobiformes - Orectolobidae	"Squatirhina americana n. sp." = Restesia americana⁹ "Squatirhina americana n. sp." = Restesia americana⁹ Estes 1964 wobbegong UCMP 55820
Hybodontiformes - Lonchidiidae	Lonchidion selachos n. gen. n. sp.⁹ Lonchidion selachos n. gen. n. sp.⁹ Estes 1964 elasmobranch UCMP 53898-53903

¹Brodkorb 1963; ²Carpenter 1982; ³Case et al. 2004; ⁴Clemens 1966; ⁵Clemens 1973; ⁶Database 2005; ⁷Elzanowski et al. 2001; ⁸Elzanowski and Stidham 2011; ⁹Estes 1964; ¹⁰Estes 1969; ¹¹Estes 1981; ¹²Estes et al. 1969; ¹³Estes and Sanchíz 1982; ¹⁴Fox 1974; ¹⁵Gaffney 1972; ¹⁶Gao and Fox 1996; ¹⁷Gardner 2000; ¹⁸Gardner 2008; ¹⁹Holroyd and Hutchison 2002; ²⁰Hope 2002; ²¹Krause 1992; ²²Larson and Currie 2013; ²³Longrich et al. 2012; ²⁴Longrich et al. 2012; ²⁵Lyson and Joyce 2009; ²⁶Novacek and Clemens 1977; ²⁷Sankey 2008