Barranca south of Lago Colhue-Huapi, Gran Barranca (Eocene of Argentina)

Where: Chubut, Argentina (45.7° S, 68.7° W: paleocoordinates 48.0° S, 59.3° W)

• coordinate estimated from map

• small collection-level geographic resolution

When: Barrancan mammal zone, Sarmiento Formation, Bartonian (41.3 - 38.0 Ma)

• Lowest fossil horizon, Casamayor Formation - now referred to as Sarmiento Formation

•Initially referred to the Casamayoran SALMA, which was later subdivided. Now considered type sequence for Barrancan SALMA (see Woodburne et al. 2014)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; claystone

• Bentonite and tuffs

Size classes: macrofossils, mesofossils

Collected by C. Ameghino, C. S. Williams, H. Herrera, W. Stein, Scarritt Expedition, G. F. Sternberg, L. G. Marshall, O. Gutierrez; reposited in the AMNH, FMNH

Primary reference: F. Ameghino. 1897. Mammiféres crétacés de l’Argentine (Deuxième contribution à la connaissance de la fauna mammalogique de couches à Pyrotherium) [Cretaceous mammals of Argentina (second contribution to the knowledge of the mammalian fauna of the Pyrotherium Beds)]. Boletin Instituto Geografico Argentino 18(4–9):406-521 [M. Uhen/M. Uhen/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 13779: authorized by John Alroy, entered by Jill Wertheim on 25.04.2002, edited by Richard Butler, Jelle Zijlstra, Terri Cleary, Grace Varnham, Miranta Kouvari, Franco Aspromonte and Matthew Carrano

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Mammalia
 Theriamorpha -
Ideodelphys microscopicus n. gen. n. sp.3
Ideodelphys microscopicus n. gen. n. sp.3 Ameghino 1902 marsupial
MACN 10344
 Paucituberculata - Caenolestidae
Progarzonia notostylopense n. gen. n. sp.4
Progarzonia notostylopense n. gen. n. sp.4 Ameghino 1904 marsupial
MACN A55-14, left dentary fragment with premolar. Simpson (1948) could not relocate it in the Ameghino Collection, but Simpson (1967) recorded that it was found again.
 Theriamorpha - Derorhynchidae
Coona gaudryi n. sp.15
Coona gaudryi n. sp.15 Simpson 1964 marsupial
MNHN, Tournouër Collection No. 1, two lower jaw fragments, one with right m1-3 one with left p4 and m2
 Theriamorpha - Peradectidae
Caroloameghinia mater14
Caroloameghinia mater14 Ameghino 1901 marsupial
MACN 10348, mandible with p2-3 m1-4, neotype of Simpson (1948), withdrawn by Simpson (1967) after the original holotype was relocated. AMNH 28441a and b, isolated upper molars.
 Sparassodonta -
Pseudocladosictis determinabile n. sp.3, Patene coluapiensis n. gen. n. sp.13
Pseudocladosictis determinabile n. sp.3 Ameghino 1902 metatherian
MACN 10325, premolar, holotype
Patene coluapiensis n. gen. n. sp.13 Simpson 1935 metatherian
 Sparassodonta - Proborhyaenidae
"Arminiheringia ? auceta" = Arminiheringia auceta8, "Dilestes dilobus n. gen. n. sp." = Arminiheringia auceta3, "Arminiheringia cultrata n. sp." = Arminiheringia auceta3, "Arminiheringia contigua n. sp." = Arminiheringia auceta4
"Arminiheringia ? auceta" = Arminiheringia auceta8 Ameghino 1902 metatherian
"Dilestes dilobus n. gen. n. sp." = Arminiheringia auceta3 Ameghino 1902 metatherian
MACN 10324, a complete lower molar, a broken one, and a fragment (syntypes)
"Arminiheringia cultrata n. sp." = Arminiheringia auceta3 Ameghino 1902 metatherian
MACN 10329, isolated M3
"Arminiheringia contigua n. sp." = Arminiheringia auceta4 Ameghino 1902 metatherian
MACN 10317, symphysis with right c-m1
 Sparassodonta - Borhyaenidae
Angelocabrerus daptes n. gen. n. sp.17
Angelocabrerus daptes n. gen. n. sp.17 Simpson 1970 metatherian
MMP 967M, dentary m2-4 and isolated teeth. "Surface find high in the beds", perhaps post-Casamayoran.
 Polydolopimorphia - Rosendolopidae
Rosendolops primigenium n. gen. n. sp.10
Rosendolops primigenium n. gen. n. sp.10 Goin and Candela 1996 metatherian
MLP 77-VI-14-3, ?M2, holotype
 Polydolopimorphia - Polydolopidae
Polydolopidae "gen. et sp. indet. 1"7, Polydolops crassus n. sp.3, "Polydolops thomasi n. gen. n. sp." = Polydolops thomasi, "Polydolops simplex n. sp." = Polydolops thomasi4, "Polydolops fur n. sp." = Polydolops thomasi3, Pliodolops primulus n. gen. n. sp.3, "Polydolops serra" = Polydolops serra14, "Anissodolops serrifer n. gen. n. sp." = Polydolops serra3, "Eudolops tetragonus n. gen. n. sp." = Eudolops tetragonus, "Propolymastodon cardatus n. sp." = Eudolops tetragonus11, "Promysops acuminatus n. gen. n. sp." = Eudolops tetragonus3, "Propolymastodon caroliameghinoi n. gen. n. sp." = Eudolops tetragonus11, Amphidolops serrula n. gen. n. sp.3, Pseudolops princeps n. gen. n. sp.3
Polydolopidae "gen. et sp. indet. 1"7 Ameghino 1897 metatherian
MLP 77-VI-14–5, a right maxillary fragment with P2-M2 (Fig. 10A); AMNH 28437, an isolated left M3
Polydolops crassus n. sp.3 Ameghino 1902 metatherian
MACN 10349
"Polydolops thomasi n. gen. n. sp." = Polydolops thomasi Ameghino 1897 metatherian
MACN 10338 (lectotype). AC 3171, AMNH 28434, 28440, 28449, FMNH P14717, MACN 10332 (c, e-g), 18469, 18472, MLP 77-VI-13-9, 77-VI-14-7, 77-VI-14-8, MNHN No. 3. From Simpson (1948:59): AMNH 28434 is the holotype of _Polydolops thomasi paahi_. Referred specimens of _P. t. paahi_: CNHM P14717, AMNH 28443, AMNH 28934
"Polydolops simplex n. sp." = Polydolops thomasi4 Ameghino 1897 metatherian
MACN 10335
"Polydolops fur n. sp." = Polydolops thomasi3 Ameghino 1897 metatherian
MACN 10342
Pliodolops primulus n. gen. n. sp.3 Ameghino 1902 metatherian
MACN 10353
"Polydolops serra" = Polydolops serra14 Ameghino 1902 metatherian
MACN 10361, mandible m2-3; MACN 10363, mandible m4 and partial m3
"Anissodolops serrifer n. gen. n. sp." = Polydolops serra3 Ameghino 1902 metatherian
MACN 10359
"Eudolops tetragonus n. gen. n. sp." = Eudolops tetragonus Ameghino 1897 metatherian
MACN 10358, an isolated right M1 missing posterolabial corner (holotype). AMNH 28437, MACN 10340b, MLP 77-VI-14-5
"Propolymastodon cardatus n. sp." = Eudolops tetragonus11 Ameghino 1897 metatherian
MACN 10333, part of a left mandibular ramus with M1-2
"Promysops acuminatus n. gen. n. sp." = Eudolops tetragonus3 Ameghino 1897 metatherian
MACN 10340a, anterior part of an edentulous right mandibular ramus with alveoli of two small incisors and C, part of diastema and alveoli of P8, and anterior alveolus of M1
"Propolymastodon caroliameghinoi n. gen. n. sp." = Eudolops tetragonus11 Ameghino 1897 metatherian
MACN 10334, greater part of a left mandibular ramus with P3-M3 complete and an isolated right C
Amphidolops serrula n. gen. n. sp.3 Ameghino 1902 metatherian
MACN 10357, an isolated left M2 (holotype). AMNH 28438
Pseudolops princeps n. gen. n. sp.3 Ameghino 1902 metatherian
MACN 10332a, an isolated left P3 (holotype). MACN 10332b-g
 Notoungulata - Archaeohyracidae
Eohyrax praerusticus n. sp.3, Eohyrax isotemnoides n. sp.3
Eohyrax praerusticus n. sp.3 Ameghino 1902 notoungulate
MACN 10780, m?1. Other syntypes are not of this species.
Eohyrax isotemnoides n. sp.3 Ameghino 1904 notoungulate
MACN 10776, dentary p4-m3 (type), upper molar, m3. MACN 10778, M1 or M2, m3, dentary with m3 and partial m2. AMNH 28665, dentary p1-m3. AMNH 28628, right dentary p1-m3, left p4-m2. AMNH 28844, M1-2.
 Notoungulata - Isotemnidae
"Plexotemnus complicatissimus n. sp." = Acoelohyrax complicatissimus4, "Dimerostephanos colhuehuapensis n. sp." = Isotemnus colhuehuapiensis3, "Isotemnus primitivus" = Isotemnus primitivus16, "Amphitemnus transitorius n. sp." = Isotemnus primitivus4, "Trimerostephanos angustus n. sp." = Isotemnus primitivus, "Amphitemnus nucleatus n. sp." = Isotemnus primitivus4, "Isotemnus conspiquus n. sp." = Isotemnus primitivus, "Tychostylops simus n. sp." = Pleurostylodon similis4, "Porotemnus crassiramis n. sp." = Pleurostylodon crassiramis3, "Pleurostylodon modicus" = Pleurostylodon modicus15, "Anchistrum sulcosum n. sp." = Pleurostylodon modicus2, "Dimerostephanos attritus n. sp." = Pleurostylodon modicus3, "Paratemnus geminatus n. sp." = Pleurostylodon modicus4, "Trimerostephanos biconus n. sp." = Pleurostylodon modicus, "Tychostylops marculus n. sp." = Pleurostylodon modicus2, "Pleurostylodon divisus n. sp." = Pleurostylodon modicus2, Thomashuxleya rostrata15, Coelostylodon florentinoameghinoi n. gen. n. sp.17, Coelostylodon caroloameghinoi n. sp.17, Anisotemnus distentus15
"Plexotemnus complicatissimus n. sp." = Acoelohyrax complicatissimus4 Ameghino 1904 notoungulate
MACN A55-1, upper and lower jaw fragments apparently of the same individual (type)
"Dimerostephanos colhuehuapensis n. sp." = Isotemnus colhuehuapiensis3 Ameghino 1902 notoungulate
MACN 10604, dentary p4-m3 (lectotype)
"Isotemnus primitivus" = Isotemnus primitivus16 Ameghino 1897 notoungulate
"Amphitemnus transitorius n. sp." = Isotemnus primitivus4 Ameghino 1897 notoungulate
MACN 10623, m2-3
"Trimerostephanos angustus n. sp." = Isotemnus primitivus Ameghino 1897 notoungulate
MACN 10593, dentary p2-m2
"Amphitemnus nucleatus n. sp." = Isotemnus primitivus4 Ameghino 1897 notoungulate
MACN 10619, broken left m3
"Isotemnus conspiquus n. sp." = Isotemnus primitivus Ameghino 1897 notoungulate
MACN 10583, maxilla M1-2 (lectotype), left dentary m2-3
"Tychostylops simus n. sp." = Pleurostylodon similis4 Ameghino 1901 notoungulate
MACN 10607, right P4 and M2-3, left P4, M1, M3
"Porotemnus crassiramis n. sp." = Pleurostylodon crassiramis3 Ameghino 1902 notoungulate
MACN 10610, dentary with p4 (lectotype), possibly associated m1 and p3, and another lower premolar
"Pleurostylodon modicus" = Pleurostylodon modicus15 Ameghino 1897 notoungulate
Types of synonyms, and: MACN 10554, almost complete lower jaw; AMNH 28878, partial skull; AMNH 28644, left P1-m3 and right M1-3; AMNH 28763, dentary p4-m2; AMNH 28630, dentary p2-m3; AMNH 28689, dentary p3-m3; AMNH 28717, jaws with right P3-m3, right p1-2, left p1-4; AMNH 28709, m1-3; AMNH 28948, dentary p3-m3; AMNH 28880, partial skull; AMNH 28646, partial skull; CNHM P13528, skull; CNHM P13296, partial skull; CNHM P13309, partial skull; CNHM P13620, maxilla with dP2-4, M1-3, P2-4
"Anchistrum sulcosum n. sp." = Pleurostylodon modicus2 Ameghino 1897 notoungulate
MACN 10767, maxilla dP2-4, M1
"Dimerostephanos attritus n. sp." = Pleurostylodon modicus3 Ameghino 1897 notoungulate
MACN 10609, dentary with parts of m2-3
"Paratemnus geminatus n. sp." = Pleurostylodon modicus4 Ameghino 1897 notoungulate
MACN 10606, maxilla P1-M2
"Trimerostephanos biconus n. sp." = Pleurostylodon modicus Ameghino 1897 notoungulate
MACN 10548, dentary m1-2 (lectotype), edentulous dentary, m2, astragalus, calcaneum
"Tychostylops marculus n. sp." = Pleurostylodon modicus2 Ameghino 1897 notoungulate
MACN 10625, left M3, type
"Pleurostylodon divisus n. sp." = Pleurostylodon modicus2 Ameghino 1897 notoungulate
MACN 10567, broken M?2 (lectotype), three premolars
Thomashuxleya rostrata15 Ameghino 1901 notoungulate
MACN 10542, M1-3; AMNH 28692, lower jaws; AMNH 28764, dentary p4
Coelostylodon florentinoameghinoi n. gen. n. sp.17 Simpson 1970 notoungulate
MMP 723M, skull
Coelostylodon caroloameghinoi n. sp.17 Simpson 1970 notoungulate
MACN 10753, partial skull with right C, P2-M3; left I1-3, C, P2-M3 (neotype)
Anisotemnus distentus15 Ameghino 1901 notoungulate
AMNH 28696, mandible with right i2-m3 and left p3-m2; AMNH 29389, dentary p3-m3
 Notoungulata - Oldfieldthomasiidae
Paginula parca8, "Oldfieldthomasia debilitata" = Oldfieldthomasia debilitata15, "Oldfieldthomasia transversa n. sp." = Oldfieldthomasia debilitata2, "Oldfieldthomasia cuneata n. sp." = Oldfieldthomasia debilitata2, "Oldfieldthomasia plicata n. sp." = Oldfieldthomasia debilitata4, "Antepithecus innexus n. sp." = Oldfieldthomasia debilitata4, "Oldfieldthomasia furcata n. sp." = Oldfieldthomasia debilitata2, "Oldfieldthomasia cingulata n. sp." = Oldfieldthomasia debilitata2, "Oldfieldthomasia conifera n. sp." = Oldfieldthomasia debilitata2, "Oldfieldthomasia parvidens n. sp." = Oldfieldthomasia parvidens2, "Oldfieldthomasia pulchella n. sp." = Oldfieldthomasia parvidens2, "Eostylops obliquatus n. sp." = Maxschlosseria praeterita2, Ultrapithecus rutilans n. gen. n. sp.2
Paginula parca8 Ameghino 1901 notoungulate
"Oldfieldthomasia debilitata" = Oldfieldthomasia debilitata15 Ameghino 1901 notoungulate
Types of synonyms; and: MACN 10765, partial skull; MACN 10761, P4-M3; MACN 10749, four dentary fragments; MACN 10750, three dentary fragments; AMNH 28680, associated dentitions; AMNH 28780, maxila I2-P3 and endocranial cast; AMNH 28632, partial skull; AMNH 28691, partial skull; AMNH 28896, partial skull; AMNH 28730, mandible left p1-m2, right i1-c1; AMNH 28963, mandible left p3-m3, right p2-m2
"Oldfieldthomasia transversa n. sp." = Oldfieldthomasia debilitata2 Ameghino 1901 notoungulate
MACN 10754, isolated upper teeth, probably dP4 and M1 (type)
"Oldfieldthomasia cuneata n. sp." = Oldfieldthomasia debilitata2 Ameghino 1901 notoungulate
MACN 10762, maxilla P1-M3
"Oldfieldthomasia plicata n. sp." = Oldfieldthomasia debilitata4 Ameghino 1901 notoungulate
MACN 10764, right dM4 (type or lectotype), left M2-3, broken M1
"Antepithecus innexus n. sp." = Oldfieldthomasia debilitata4 Ameghino 1901 notoungulate
MACN 10839, isolated upper molar
"Oldfieldthomasia furcata n. sp." = Oldfieldthomasia debilitata2 Ameghino 1901 notoungulate
MACN 10748, left maxilla with P1-M3
"Oldfieldthomasia cingulata n. sp." = Oldfieldthomasia debilitata2 Ameghino 1901 notoungulate
MACN 10772, skull
"Oldfieldthomasia conifera n. sp." = Oldfieldthomasia debilitata2 Ameghino 1901 notoungulate
MACN 10757, maxilla P4-M3
"Oldfieldthomasia parvidens n. sp." = Oldfieldthomasia parvidens2 Ameghino 1901 notoungulate
MACN 10763, right P4-M2 (type) and three isolated upper cheek teeth; MHN Tournouër 21, partial skull
"Oldfieldthomasia pulchella n. sp." = Oldfieldthomasia parvidens2 Ameghino 1901 notoungulate
MACN 10756, maxilla fragment with dP3-4 and M1
"Eostylops obliquatus n. sp." = Maxschlosseria praeterita2 Ameghino 1901 notoungulate
MACN 10482, maxilla M2
Ultrapithecus rutilans n. gen. n. sp.2 Ameghino 1901 notoungulate
MACN 10818, maxilla P1-M2 (type); AMNH 28706, palate P1-M3; AMNH 28583, palate with left dI2-dP4 and M1, right dP1-4, M1, part of M2; MHN Tournouër No. 23, maxilla P2-M3
 Notoungulata -
Tonostylops spissus n. gen. n. sp.3
Tonostylops spissus n. gen. n. sp.3 Ameghino 1902 notoungulate
MACN 10483, dentary ?m2-3
 Notoungulata - Henricosborniidae
Henricosbornia ? lophodonta8
Henricosbornia ? lophodonta8 Ameghino 1901 notoungulate
 Notoungulata - Archaeopithecidae
"Archaeopithecus rogeri n. gen. n. sp." = Archaeopithecus rogeri, "Adpithecus plenus n. sp." = Archaeopithecus rogeri3, "Notopithecus fossulatus n. sp." = Archaeopithecus rogeri
"Archaeopithecus rogeri n. gen. n. sp." = Archaeopithecus rogeri Ameghino 1897 notoungulate
MACN 10816, maxilla P1-M2
"Adpithecus plenus n. sp." = Archaeopithecus rogeri3 Ameghino 1897 notoungulate
MACN 10851, partial M1 and complete M2
"Notopithecus fossulatus n. sp." = Archaeopithecus rogeri Ameghino 1897 notoungulate
MACN 10824, dentary p3-4 (lectotype); unassociated dentary with p4-m1; P?2
 Notoungulata - Notostylopidae
Notostylops bicinctus n. sp., "Notostylops murinus n. gen. n. sp." = Notostylops murinus, "Notostylops promurinus n. sp." = Notostylops murinus4, "Notostylops chicoensis n. sp." = Notostylops murinus3, "Entelostylops completus n. sp." = Notostylops murinus2, "Isostylops fretus n. sp." = Notostylops murinus3, Catastylops deflexus n. sp.4, "Notostylops parvus n. sp." = Homalostylops parvus, "Homalostylops rigeo n. sp." = Homalostylops parvus2, "Acrostylops pungiunculus n. sp." = Homalostylops parvus2, "Entelostylops incolumis n. sp." = Homalostylops parvus2
Notostylops bicinctus n. sp. Ameghino 1897 notoungulate
MACN 10512, left upper premolar
"Notostylops murinus n. gen. n. sp." = Notostylops murinus Ameghino 1897 notoungulate
MACN 10500, partial skull (paralectotype); MACN 10498, both sides of lower jaw (lectotype). Numerous referred specimens.
"Notostylops promurinus n. sp." = Notostylops murinus4 Ameghino 1897 notoungulate
MACN 10507, maxilla with P3-M3, molars broken
"Notostylops chicoensis n. sp." = Notostylops murinus3 Ameghino 1897 notoungulate
MACN 10518, right m1-2
"Entelostylops completus n. sp." = Notostylops murinus2 Ameghino 1897 notoungulate
MACN 10492, maxilla P4-m3
"Isostylops fretus n. sp." = Notostylops murinus3 Ameghino 1897 notoungulate
MACN 10494, four isolated premolars
Catastylops deflexus n. sp.4 Ameghino 1904 notoungulate
MACN 10493, maxilla with P2-4
"Notostylops parvus n. sp." = Homalostylops parvus Ameghino 1897 notoungulate
MACN 10527, right p2-m3, left p3-m2; MACN 10534, left p3-m3; probably all one individual
"Homalostylops rigeo n. sp." = Homalostylops parvus2 Ameghino 1897 notoungulate
MACN 10473, partial lower jaw with left p3-m3, right p3-4
"Acrostylops pungiunculus n. sp." = Homalostylops parvus2 Ameghino 1897 notoungulate
MACN 10468, mandible with right i3 and p1-m2, left i2 and p3-4
"Entelostylops incolumis n. sp." = Homalostylops parvus2 Ameghino 1897 notoungulate
MACN 10495, four uppre molars
 Notoungulata -
"Acoelodus" proclivus n. sp.3
"Acoelodus" proclivus n. sp.3 Ameghino 1902 notoungulate
MACN 10752, isolated M1 (type), jaw fragment with p3-4
 Notoungulata - Notopithecidae
"Antepithecus brachystephanus n. sp." = Antepithecus brachystephanus2, "Antepithecus interrasus n. sp." = Antepithecus brachystephanus4, "Infrapithecus cinctus n. sp." = Antepithecus brachystephanus2, Transpithecus obtentus n. sp.2, "Notopithecus adapinus n. gen. n. sp." = Notopithecus adapinus, "Adpithecus subtenuis n. sp." = Notopithecus adapinus3, "Adpithecus secans" = Notopithecus adapinus15
"Antepithecus brachystephanus n. sp." = Antepithecus brachystephanus2 Ameghino 1901 notoungulate
MACN 10841, seven isolated upper teeth (including lectotype M2), fragmentary maxilla, two fragmentary dentaries
"Antepithecus interrasus n. sp." = Antepithecus brachystephanus4 Ameghino 1901 notoungulate
MACN 10859, maxillary P2-M2
"Infrapithecus cinctus n. sp." = Antepithecus brachystephanus2 Ameghino 1901 notoungulate
MACN 10826, dentary p2-m3
Transpithecus obtentus n. sp.2 Ameghino 1901 notoungulate
MACN-A 10833a, maxilla M1-3 (lectotype). AMNH 28791, P2-P4; AMNH 28861, dentary p2-m3; AMNH 28953, maxilla P3-M3; FMNH P14895, dentary m3; MACN-A 10824, P2; MACN-A 10833b, M2; MACN-A 10833c, M3; MACN-A 10845, P4; MPEF-PV 1284, dentary p3-m1 (from "Puesto Almendra")
"Notopithecus adapinus n. gen. n. sp." = Notopithecus adapinus Ameghino 1897 notoungulate
Syntypes: MACN 10822, associated jaw, limb bone, and vertebra fragments; and MACN 10843, jaw fragments and isolated teeth.
"Adpithecus subtenuis n. sp." = Notopithecus adapinus3 Ameghino 1897 notoungulate
MACN 10834, maxilla P3-M3 (lectotype), maxilla P4-M2
"Adpithecus secans" = Notopithecus adapinus15 Ameghino 1897 notoungulate
Referred specimens in Ameghino Collection.
 Astrapotheria - Trigonostylopidae
"Albertogaudrya unica" = Albertogaudrya unica15, "Scabellia laticincta n. sp." = Albertogaudrya unica2, "Albertogaudrya tersa n. sp." = Albertogaudrya unica3, "Albertogaudrya oxygona n. sp." = Albertogaudrya unica4, "Albertogaudrya separata n. sp." = Albertogaudrya unica4, "Scabellia cyclogona n. sp." = Albertogaudrya unica4, ? Trigonostylops duplex n. sp.2, "Trigonostylops wortmani n. gen. n. sp." = Trigonostylops wortmani, "Trigonostylops insumptus n. sp." = Trigonostylops wortmani2, "Trigonostylops scabellum" = Trigonostylops wortmani15, "Trigonostylops columnifer n. sp." = Trigonostylops wortmani4, "Trigonostylops eximius n. sp." = Trigonostylops wortmani2, "Trigonostylops germinalis n. sp." = Trigonostylops wortmani4, "Trigonostylops hemicyclus n. sp." = Trigonostylops wortmani2, "Trigonostylops secundarius n. sp." = Trigonostylops wortmani2, "Trigonostylops minimus n. sp." = Trigonostylops wortmani2, "Trigonostylops trigonus n. sp." = Trigonostylops wortmani2, "Trigonostylops coryphodontoides n. sp." = Trigonostylops wortmani4
"Albertogaudrya unica" = Albertogaudrya unica15 Ameghino 1901 placental
AMNH 28639, P4-M1
"Scabellia laticincta n. sp." = Albertogaudrya unica2 Ameghino 1901 placental
MACN 12008, partial upper cheek tooth
"Albertogaudrya tersa n. sp." = Albertogaudrya unica3 Ameghino 1901 placental
MACN 16033, upper canine; MACN 12012, incisor. Type apparently an upper molar, not relocated by Simpson (1967).
"Albertogaudrya oxygona n. sp." = Albertogaudrya unica4 Ameghino 1901 placental
MACN 12007, partial upper molar
"Albertogaudrya separata n. sp." = Albertogaudrya unica4 Ameghino 1901 placental
MACN 12004, upper molar
"Scabellia cyclogona n. sp." = Albertogaudrya unica4 Ameghino 1901 placental
MACN 12005, upper cheek tooth fragments
? Trigonostylops duplex n. sp.2 Ameghino 1901 placental
MACN 12010, broken symphysis
"Trigonostylops wortmani n. gen. n. sp." = Trigonostylops wortmani Ameghino 1897 placental
MACN 10651, M?2, lectotype, P?4, m3 talonid (type series). Other specimens including AMNH 28700, skull
"Trigonostylops insumptus n. sp." = Trigonostylops wortmani2 Ameghino 1897 placental
MACN 10654, M?2
"Trigonostylops scabellum" = Trigonostylops wortmani15 Ameghino 1897 placental
MACn 10629, p2-4
"Trigonostylops columnifer n. sp." = Trigonostylops wortmani4 Ameghino 1897 placental
MACN 10638, two upper molars including the lectotype
"Trigonostylops eximius n. sp." = Trigonostylops wortmani2 Ameghino 1897 placental
MACN 10650, upper molar (lectotype), two m3s, m3 talonid, m?2, m?1, C1
"Trigonostylops germinalis n. sp." = Trigonostylops wortmani4 Ameghino 1897 placental
MACN 10639, upper molar
"Trigonostylops hemicyclus n. sp." = Trigonostylops wortmani2 Ameghino 1897 placental
MACN 10659, m2, lectotype, and another m2
"Trigonostylops secundarius n. sp." = Trigonostylops wortmani2 Ameghino 1897 placental
MACN 10645, upper molar (lectotype), upper molar, upper premolar
"Trigonostylops minimus n. sp." = Trigonostylops wortmani2 Ameghino 1897 placental
MACN 10660, M?3
"Trigonostylops trigonus n. sp." = Trigonostylops wortmani2 Ameghino 1897 placental
MACN 10656, broken upper molar
"Trigonostylops coryphodontoides n. sp." = Trigonostylops wortmani4 Ameghino 1897 placental
MACN 10641, upper molar
 Cingulata - Peltephilidae
Peltephilidae indet.6
 Cingulata - Dasypodidae
"Anteutatus lenis" = Utaetus lenis14, Utaetus deustus n. sp.3, "Utaetus buccatus n. sp." = Utaetus buccatus3, "Orthutaetus clavatus n. sp." = Utaetus buccatus3, "Posteutatus indentatus n. sp." = Utaetus buccatus3, "Posteutatus indemnis n. sp." = Utaetus buccatus3, "Orthutaetus crenulatus n. sp." = Utaetus buccatus3, "Posteutatus scabridus n. sp." = Utaetus buccatus3, Prostegotherium notostylopianum n. sp.3, Prostegotherium astrifer n. sp.3, Parutaetus signatus n. sp.3, Parutaetus clusus n. sp.3, Parutaetus chicoensis n. sp.3, Meteutatus percarinatus n. sp.3, Machlydotherium sp.6, "? Machlydotherium sparsus n. sp." = Machlydotherium sparsum3, "Pseudostegotherium" chubutanum n. sp.3
"Anteutatus lenis" = Utaetus lenis14 Ameghino 1902 armadillo
MACN 10459, five scutes. Holotype may also be from here.
Utaetus deustus n. sp.3 Ameghino 1902 armadillo
MACN 10431, four scutes
"Utaetus buccatus n. sp." = Utaetus buccatus3 Ameghino 1902 armadillo
AMNH 28668, neotype, 200-300 scutes; part of cephalic shield, two areas of associated buckler scutes, two areas of movable bands, connected parietals with parts of squamosals and temporals, fragments of occipitals, crushed maxilla with four teeth, both sides of lower jaw with most of the teeth, nine isolated teeth; parts of nine vertebrae; one chevron bone, several rib fragments; parts of both scapulae, one humerus and ulna, and both radii; one ilium and fragment of the other, parts of one ischium and pubis, parts of one femur and fibula, and various tarsals, metatarsals, and phalanges. Original holotype was MACN 10457, 18 scutes, metacarpal, and calcaneum, locality uncertain.
"Orthutaetus clavatus n. sp." = Utaetus buccatus3 Ameghino 1902 armadillo
MACN 10440, scute
"Posteutatus indentatus n. sp." = Utaetus buccatus3 Ameghino 1902 armadillo
MACN 10464, three scutes
"Posteutatus indemnis n. sp." = Utaetus buccatus3 Ameghino 1902 armadillo
MACN 10442, eight scutes
"Orthutaetus crenulatus n. sp." = Utaetus buccatus3 Ameghino 1902 armadillo
MACN 10430, scute
"Posteutatus scabridus n. sp." = Utaetus buccatus3 Ameghino 1902 armadillo
MACN 10463, three scutes
Prostegotherium notostylopianum n. sp.3 Ameghino 1902 armadillo
MACN 10451, 33 scutes
Prostegotherium astrifer n. sp.3 Ameghino 1902 armadillo
MACN 10444, 18 scutes
Parutaetus signatus n. sp.3 Ameghino 1902 armadillo
MACN 10422, six scutes
Parutaetus clusus n. sp.3 Ameghino 1902 armadillo
MACN 10436, two scutes
Parutaetus chicoensis n. sp.3 Ameghino 1902 armadillo
MACN 10429, six scutes
Meteutatus percarinatus n. sp.3 Ameghino 1902 armadillo
MACN 10453, buckler scute
Machlydotherium sp.6 Ameghino 1902 armadillo
"? Machlydotherium sparsus n. sp." = Machlydotherium sparsum3 Ameghino 1902 armadillo
MACN 10427, two broken buckler scutes. Simpson (1948) suspects these specimens were really from the Mustersan. They were said to be from the upper part of the "Notostylopéen" (?= El Nuevo level).
"Pseudostegotherium" chubutanum n. sp.3 Ameghino 1902 armadillo
MACN A-10438 (carapace osteoderms, syntype); MPEF PV 5425, 6344, 6398, 6465, 6516, 7234A, 7660B; AMNH 28745, 28851
 Placentalia -
Pilosa indet.6
Pilosa indet.6 edentate
 Panameriungulata - Didolodontidae
"Lambdaconus mamma n. sp." = Paulogervaisia mamma2, "Lambdaconus porcus n. sp." = Paulogervaisia porca2, Paulogervaisia inusta n. sp.2, Didolodus minor9, "Didolodus multicuspis" = Didolodus multicuspis14, "Lonchoconus lanceolatus n. gen. n. sp." = Didolodus multicuspis2, "Didolodus colligatus n. sp." = Didolodus multicuspis3, Proectocion precisus n. sp.4, Proectocion argentinus n. sp.4, Ernestokokenia nitida14
"Lambdaconus mamma n. sp." = Paulogervaisia mamma2 Ameghino 1901 placental
MACN 10719: fragmentary mandible p3-m2; maxilla with broken M2-3; broken M2; other fragments. Simpson (1948:107) selected the maxilla as the lectotype and doubted the referral of the other fragments.
"Lambdaconus porcus n. sp." = Paulogervaisia porca2 Ameghino 1901 placental
MACN 10721, maxilla fragment with broken M2 and complete M3
Paulogervaisia inusta n. sp.2 Ameghino 1901 placental
MACN 10664, right m3, type of Paulogervaisia inusta; MACN 10719, right jaw fragment with p3-m2 and maxillary fragment with M2-3 and isolated right M2, type of Lambdaconus mamma; MPEF 7621, isolated right m2.
Didolodus minor9 Simpson 1948 placental
"Didolodus multicuspis" = Didolodus multicuspis14 Ameghino 1897 placental
MACN 10720, mandible with m3 and possibly associated isolated M1 and C; MACN 10738, right P1 and left P2; MACN 10729, left M2; AMNH 28475, mandible p4, m2-3; AMNH 2878, M3; AMNH 28774, p4. Additional specimens are types of synonyms.
"Lonchoconus lanceolatus n. gen. n. sp." = Didolodus multicuspis2 Ameghino 1897 placental
MACN 10730, left M1
"Didolodus colligatus n. sp." = Didolodus multicuspis3 Ameghino 1897 placental
MACN 10736, right and left m2, left p3-4
Proectocion precisus n. sp.4 Ameghino 1904 placental
MACN 10679, maxilla with P4-M3
Proectocion argentinus n. sp.4 Ameghino 1904 placental
MACN 10673, M3
Ernestokokenia nitida14 Ameghino 1901 placental
AMNH 28477, left m3
 Panameriungulata - Proterotheriidae
Ricardolydekkeria praerupta n. sp.2, Ricardolydekkeria profunda n. sp.2, Guilielmofloweria plicata n. gen. n. sp.2, Anisolambda fissidens8, Josepholeidya deculca n. sp.2
Ricardolydekkeria praerupta n. sp.2 Ameghino 1901 placental
MACN 10701, right upper cheek tooth
Ricardolydekkeria profunda n. sp.2 Ameghino 1901 placental
MACN 10706, partial right upper cheek tooth (lectotype); two fragments of upper teeth and two of lower molars.
Guilielmofloweria plicata n. gen. n. sp.2 Ameghino 1901 placental
MACN 10703, left upper cheek tooth (type)
Anisolambda fissidens8 Ameghino 1901 placental
Josepholeidya deculca n. sp.2 Ameghino 1901 placental
MACN 10682, upper molar
 Panameriungulata - Macraucheniidae
Rutimeyeria conulifera n. gen. n. sp.2
Rutimeyeria conulifera n. gen. n. sp.2 Ameghino 1901 placental
MACN 12013, upper molar
  -
Proplanodus adnepos n. gen. n. sp.3
Proplanodus adnepos n. gen. n. sp.3 Ameghino 1902 mammal
MACN 12015, four ?incisors
Reptilia
 Squamata - Madtsoiidae
? Madtsoia sp.1
? Madtsoia sp.1 Simpson 1933 snake
MACN-A 10879a, MACN-A 10879b, MACN-A 10888, three fragments of trunk vertebrae's centra; MACN-A 10892a, centrum fragment with right prezyg. of a trunk vertebra; MACN-A 10892b, centrum fragment with left prezyg. of a trunk vertebra; MACN-A 10873a and 10873b, fragments of a trunk vertebrae.
 Squamata - Boidae
? Boa sp.1
? Boa sp.1 Linnaeus 1758 boa
MACN-A 10875, partial trunk vertebra
 Loricata -
Crocodylomorpha indet.12
Crocodylomorpha indet.12 crocodilian
MACN 10871, tooth
Actinopteri
 Amiiformes - Amiidae
Pappichthys patagonica n. gen. n. sp.5
Pappichthys patagonica n. gen. n. sp.5 bowfin
MACN A-11263, anterior part of a left dentary (holotype). MACN A-11592, abdominal and caudal centra, and an indeterminate bone scrap
1Albino 1993; 2Ameghino 1901; 3Ameghino 1902; 4Ameghino 1904; 5Ameghino 1906; 6Carlini et al. 2005; 7Chornogubsky 2020; 8Cifelli 1985; 9Gelfo 2010; 10Goin and Candela 1996; 11Marshall 1982; 12Simpson 1932; 13Simpson 1935; 14Simpson 1948; 15Simpson 1967; 16Simpson 1967; 17Simpson 1970