Bentiaba (layer above Bench 19) (Cretaceous of Angola)

Also known as Sao Nicolau

Where: Namibe, Angola (14.3° S, 12.4° E: paleocoordinates 25.0° S, 1.7° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Mocuio Formation, Early/Lower Maastrichtian (70.6 - 66.0 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: marginal marine; fine-grained siltstone

• fine-grained sandstone

Size class: macrofossils

Collected in 2005 - present

Primary reference: M. T. Antunes. 1964. O Neocretácico e o Cenozoico do litoral de Angola. Junta de Investigações do Ultramar 1-254 [M. Uhen/M. Uhen/M. Uhen]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106705: authorized by John Alroy, entered by John Alroy on 28.03.2011, edited by Philip Mannion and Jonathan Tennant

Creative Commons license: CC BY (attribution)

Taxonomic list

• not the same as a previous Bentiaba = San Nicolau collection reported by Antunes 1964
unclassified
  -
Mollusca spp.2 Linnaeus 1758
Gnathostomata
  -
Chondrichthyes spp.2 Huxley 1880 cartilaginous fish
Osteichthyes spp.2 bony fish
Reptilia
 Testudines -
Toxochelys sp.2 Cope 1873 turtle
MGUAN-PA168, hyoplastron, peripherals and costal plates
 Testudines - Pancheloniidae
Euclastes sp.2 Cope 1870 turtle
MGUAN-PA157, complete skull and mandible, cervical vertebrae, peripheral plates, forelimb
 Testudines - Protostegidae
cf. Calcarichelys sp.2 Zangerl 1953 sea turtle
MGUAN-PA167, neural scute
Protostega sp.2 Cope 1871 sea turtle
MGUAN-PA158, two humeri, three costal plates, one xiphiplastron, one peripheral plate (single animal)
 Squamata - Mosasauridae
Halisaurus sp.2 Marsh 1869 mosasaur
MGUAN-PA18, two partial skeletons including many elements of the skulls, vertebrae, ribs and limbs
Phosphorosaurus sp.2 Dollo 1889 mosasaur
MGUAN-PA52, isolated partial frontal
cf. Platecarpus ptychodon2 Arambourg 1952 mosasaur
MGUAN-PA160, two partial skulls and skeletons including limb material, isolated tooth crowns [new genus not erected but required]
Liodon sp.2 Owen 1842 mosasaur
Prognathodon kianda n. sp.5 Schulp et al. 2008 mosasaur
MGUAN-PA129 (holotype), MGUAN-PA128, MGUAN-PA149, MGUAN-PA150, MGUAN-PA151 - several specimens of different ontogenetic ages; shed teeth, isolated bones, partial skulls and skeletons
Carinodens belgicus4 Woodward 1891 mosasaur
MGUAN-PA 171
Globidens phosphaticus3 Bardet et al. 2005 mosasaur
PA24 (a partial skeleton including many elements of the skull and some postcrania) and PA23 (a fragmentary skull and posterior dorsal vertebra)
 Dinosauria -
Dinosauria indet.2 Owen 1842 dinosaur
MGUAN-PA175, distal part of humerus
 Ornithischia -
Hadrosauroidea indet.2 Huene 1954 ornithopod
MGUAN-PA176, a single phalanx
 Pterosauria -
Pterosauria indet.1 Kaup 1834 pterosaur
MGUAN-PA652, 653, 656-659
Pterodactyloidea indet.1 pterosaur
MGUAN-PA660
Pteranodontia indet.1 Marsh 1876 pterosaur
MGUN-PA163, left femur
Epapatelo otyikokolo n. gen. n. sp.1 Fernandes et al. 2022 pterosaur
MGUAN-PA650 - holotype; referred material: MGUAN-PA661
 Plesiosauria - Elasmosauridae
Elasmosauridae indet.2 Cope 1869 elasmosaur
MGUAN-PA113, basicranium, and MGUAN-PA126, symphyseal region of the mandibl, various unnumbered specimens, teeth
cf. Tuarangisaurus sp.2 Wiffen and Moisley 1986 elasmosaur
MGUAN-PA106