PBDB Collection

Upper Site (Miocene to of Australia)

Also known as Godthelp’s Hill, Riversleigh

Where: Queensland, Australia (19.0° S, 138.7° E: paleocoordinates 30.7° S, 134.7° E)

• coordinate stated in text

• outcrop-level geographic resolution

When: System B Formation, Aquitanian to Aquitanian (23.0 - 16.0 Ma)

• System B is "early Miocene" based on biochronology

Environment/lithology: terrestrial; limestone

Size class: mesofossils

Collection methods: acetic

• Queensland Museum collection (QM)

•Michael Archer reference collection (AR), University of New South Wales (possibly serving as temporary registration)

Primary reference: K. K. Roberts, M. Archer, S. J. Hand and H. Godthelp. 2007. New genus and species of extinct Miocene ringtail possums (Marsupialia: Pseudocheiridae). American Museum Novitates 3560:1-15 [J. Alroy/J. Alroy/M. Uhen]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106497: authorized by John Alroy, entered by John Alroy on 21.03.2011, edited by Albert Garcia Selles, Torsten Liebrecht, Matthew Clapham, Mark Uhen, Bethany Allen and Grace Varnham

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Coleoptera -	Polyphaga indet.⁷ Polyphaga indet.⁷ Emery 1886 beetle QM F34583, F34586
Coleoptera - Curculionidae	Curculionidae indet.⁷ Curculionidae indet.⁷ Latreille 1802 snout beetle QM F16648, F34585
Coleoptera - Histeridae	Histeridae indet.⁷ Histeridae indet.⁷ Gyllenhal 1808 clown beetle QM F34582
Coleoptera - Ommatidae	Ommatidae indet.⁷ Ommatidae indet.⁷ Sharp and Muir 1912 beetle QM F34595
Dicondylia -	Trichoptera indet.⁷ Trichoptera indet.⁷ Kirby 1815 caddisfly QM F34584, F34587-F34594
Myriapoda
Chilognatha -	Julida indet.⁷ Julida indet.⁷ Leach 1814 millipede QM F34596
Mammalia
Peramelemorphia -	Galadi speciosus¹³ Galadi speciosus¹³ Travouillion et al. 2010 marsupial
Peramelemorphia - Yaralidae	Yarala burchfieldi⁹ Yarala burchfieldi⁹ Muirhead and Filan 1995 marsupial
Diprotodontia - Phascolarctidae	Litokoala garyjohnstoni n. sp.⁸ Litokoala garyjohnstoni n. sp.⁸ Louys et al. 2007 koala
Diprotodontia -	Ganawamaya acris n. gen. n. sp.⁴, Balbaroo fangaroo n. gen. n. sp.⁵ Ganawamaya acris n. gen. n. sp.⁴ Cooke 1992 diprotodont marsupial Balbaroo fangaroo n. gen. n. sp.⁵ Cooke 2000 diprotodont marsupial
Diprotodontia - Miralinidae	Durudarwiri inusitatus n. gen. n. sp.⁶ Durudarwiri inusitatus n. gen. n. sp.⁶ Crosby and Archer 2000 diprotodont marsupial
Diprotodontia - Macropodidae	Gumardee webbi¹⁴, Gumardee keari¹⁴, Cookeroo hortusensis³ Gumardee webbi¹⁴ Travouillon et al. 2022 kangaroo QM F19926, left maxilla with broken P3. VVC-F Site: QM F57594, left maxilla with P2, dP3, M1–2, broken M3. Gumardee keari¹⁴ Travouillon et al. 2022 kangaroo QM F19620, left maxilla with M1–4; QM F19627, right maxilla with M4; QM F19641, right juvenile dentary with m4 in crypt; QM F19685, right dentary with m1–4; QM F19930, right dentary with m1 talonid, m2–3; QM F19948, right juvenile dentary with dp3, p3 out of its crypt, m1. Cookeroo hortusensis³ Butler et al. 2016 kangaroo
Diprotodontia - Pseudocheiridae	Paljara sp., Pildra sp., Marlu sp., Pseudochirops sp., Gawinga aranaea Paljara sp. Woodburne et al. 1987 ringtail possum species name not given Pildra sp. Woodburne et al. 1987 ringtail possum species name not given Marlu sp. Woodburne et al. 1987 ringtail possum species name not given Pseudochirops sp. Marschie 1915 green ringtail species name not given Gawinga aranaea Roberts et al. 2007 ringtail possum
Notoryctemorphia - Notoryctidae	Naraboryctes philcreaseri¹ Naraboryctes philcreaseri¹ Archer et al. 2010 marsupial
Reptilia
Squamata - Madtsoiidae	Yurlunggur sp.¹², Nanowana schrenki n. sp.¹⁰, Nanowana godthelpi¹⁰ Yurlunggur sp.¹² Scanlon 1992 snake 2 partial dentaries (QM F23064, F51380) Nanowana schrenki n. sp.¹⁰ Scanlon 1997 snake QMF 31395 (holotype), right palatine; QMF 31390, 31391, 31394, maxilla fragments Nanowana godthelpi¹⁰ Scanlon 1997 snake QMF 31389, dentary; QMF 23066, palatine; QMF 31390, maxilla fragment; QMF 23067, 31385, pterygoids
Squamata - Pythonidae	Morelia riversleighensis¹¹ Morelia riversleighensis¹¹ Smith and Plane 1985 python partial skeleton of large individual, comprising right dentary and anterior part of right compound bone (both AR l3392), left maxilla (AR l3l36), 89 partial to complete vertebrae (AR 10842), and fragments of ribs including 11 rib heads (AR 13134, 13135, 14123)
Aves
Passeriformes - Menuridae	Menura tyawanoides n. sp.² Menura tyawanoides n. sp.² Boles 1995 lyrebird

¹Archer et al. 2010; ²Boles 1995; ³Butler et al. 2016; ⁴Cooke 1992; ⁵Cooke 2000; ⁶Crosby and Archer 2000; ⁷Duncan et al. 1998; ⁸Louys et al. 2007; ⁹Muirhead and Filan 1995; ¹⁰Scanlon 1997; ¹¹Scanlon 2001; ¹²Scanlon 2005; ¹³Travouillion et al. 2010; ¹⁴Travouillon et al. 2022