Tlayua Formation (Cretaceous of Mexico)

Also known as Tepexi; Tlayua Quarry

Where: Puebla, Mexico (19.1° N, 98.2° W: paleocoordinates 17.1° N, 61.6° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Tlayua Formation, Albian (113.0 - 100.5 Ma)

• formation-level stratigraphic resolution

Environment/lithology: lagoonal or restricted shallow subtidal; lithified lime mudstone

Size class: macrofossils

Preservation: adpression

Collection methods: bulk,


Primary reference: L. Martin-Medrano and P. Garcia-Barrera. 2006. Fossil Ophiuroids of Mexico. In F. J. Vega, T. G. Nybor, M. D. C. Perrillat, M. Montellano-Ballesteros, S. R. S. Cevallos-Ferriz, S. A. Quiroz-Barroso (eds.), Topics in Geobiology 24:115-131 [L. Villier/J. Millet/M. Uhen]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 80283: authorized by Loic Villier, entered by Julien Millet on 10.04.2008, edited by Jonathan Tennant, Matthew Carrano, Richard Butler, Carrie Schweitzer, Roger Benson, Loic Villier and Sharon McMullen

Creative Commons license: CC BY (attribution)

Taxonomic list

 Isopoda - Archaeoniscidae
 Decapoda - Dorippidae
Tepexicarcinus tlayuaensis5 Feldmann et al. 1998 crab
 Decapoda - Aeglidae
Protaegla miniscula5 Feldmann et al. 1998 decapod
 Teleostei - Aspidorhynchidae
 Elopiformes - Megalopidae
Megalops sp.4 Lacépède 1803
 Elopiformes - Elopidae
Notelops sp.4 Woodward 1901
 Ellimmichthyiformes - Ellimmichthyidae
Ellimmichthys sp.2 Jordan and Gilbert 1919
 Pycnodontiformes - Pycnodontidae
 Lepisosteiformes -
Lepidotes sp.4 Agassiz 1843 gar
 Ionoscopiformes - Ophiopsidae
Teoichthys kallistos4 Applegate 1988
 Halecomorphi - Ionoscopidae
Quetzalichthys perrilliatae n. gen. n. sp.1 Alvarado-Ortega and Espinosa-Arrubarrena 2008
IGM 8592, complete specimen exposed from the right side (IGM-locality 2432); IGM 8593, IGM 8595, IGM 8596
 Coelacanthiformes - Mawsoniidae
Axelrodichthys cf. araripensis4 Maisey 1986 coelacanth
 Testudines -
Testudines indet.10 Batsch 1788 turtle
Two taxa, partial skeletons
 Pterosauria -
Ornithocheiroidea indet.12 pterosaur
Humerus, scapulocoracoids, wing-bones, femur, considered similar to Pteranodon or Nyctosaurus
 Loricata - Teleosauridae
"cf. Crocodileimus sp." = Crocodilaemus4
"cf. Crocodileimus sp." = Crocodilaemus4 Jourdan 1857 marine crocodile
 Squamata -
Huehuecuetzpalli mixtecus n. gen. n. sp.8, Tepexisaurus tepexii n. gen. n. sp.11
Huehuecuetzpalli mixtecus n. gen. n. sp.8 Reynoso 1998 squamates
IGM 4185 (loc. IGM-1971-NSM number 1, level Z/10, quadrant 16/5), 7389 (loc. IGM-1995-NSF number 2, level H, quadrant 1/5)
Tepexisaurus tepexii n. gen. n. sp.11 Reynoso and Callison 2000 squamates
IGM 7466 (holotype)
 Rhynchocephalia - Sphenodontidae
Ankylosphenodon pachyostosus n. gen. n. sp.9 Reynoso 2000 rhynchocephalian
IGM 7441-7447
 Rhynchocephalia -
Pamizinsaurus tlayuanensis n. gen. n. sp.7 Reynoso 1997 rhynchocephalian
IGM 6854 (holotype)
 Dendrochirotida - Psolidae
Parapsolus tlayuensis n. gen. n. sp.3
Parapsolus tlayuensis n. gen. n. sp.3 sea cucumber
 Dendrochirotida - Cucumariidae
Paleopentacta alencasterae n. gen. n. sp.3 Applegate et al. 2009 sea cucumber
 Ophiurida - Ophiactidae
Ophiactis applegatei6 Martin-Medrano et al. 2009 brittle star
 Bennettitales -
Zamites sp.4 Brongniart 1828
 Pinales - Podocarpaceae
Podozamites sp.3 Braun 1843
 Pinales - Araucariaceae
cf. Araucaria sp.4 de Jussieu 1789
 Pinales -
Brachyphyllum sp.3 Brongniart 1828