CDZh-17a, Dzharakuduk II, Kulbeke (Cretaceous to of Uzbekistan)

Also known as Dzarakuduk, Dzhyrakuduk

Where: Navoi, Uzbekistan (42.1° N, 62.7° E: paleocoordinates 36.8° N, 57.0° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Bissekty Formation, Middle Turonian to Middle Turonian (93.5 - 89.3 Ma)

• lower part of formation, 26 m above base

•Also described as Taykarshinskaya beds.

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size class: macrofossils

Collected by L. A. Nessov in 1977

Collection methods: quarrying, mechanical,

Primary reference: L. A. Nessov. 1981. Amfibii i reptilii v ekosistemakh Mela sredney Azii [Amphibia and reptiles in Cretaceous ecosystems of central Asia]. In I. V. Darevsky (ed.), The Problems of Herpetology. Fifth Herpetological Conference. Abstracts 91-92 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 46959: authorized by Matthew Carrano, entered by Matthew Carrano on 27.01.2005, edited by Kaitlin Maguire and Roger Benson

Creative Commons license: CC BY (attribution)

Taxonomic list

• Nessov (1984) listed turtles from "Dzharakuduk (= Kulbecke, =Bissekty) locality, central Kizylkum Desert and unamed beds adajcent to Tjulkili Top", but only gave details for CDZh-17a
Chondrichthyes
 Sclerorhynchiformes - Ptychotrygonidae
Ptychotrygon sp.7 Jaekel 1894 elasmobranch
 Myliobatiformes - Dasyatidae
Myledaphus tritus7 whiptail stingray
Actinopteri
 Ichthyodectiformes -
? Aidachar sp.9 Nessov 1981
 Amiiformes - Amiidae
Amia limosa8 Nessov 1985 bowfin
 Lepisosteiformes - Lepisosteidae
Lepisosteidae indet.9 Cuvier 1825 gar
Osteichthyes
 Salientia - Discoglossidae
Aralobatrachus robustus n. gen. n. sp.
Aralobatrachus robustus n. gen. n. sp. Nessov 1981 painted frog
ZIN, PHA No 477-7 (holotype partial maxilla), LU-N4/54 (maxilla)
 Caudata - Urodela
"Mynbulakia surgayi" = Eoscapherpeton asiaticum9
"Mynbulakia surgayi" = Eoscapherpeton asiaticum9 Nessov 1981 hellbender
Reptilia
 Testudines - Macrobaenidae
Anatolemys cf. maximus6 Khozatskii and Nessov 1979 turtle
 Testudines - Trionychidae
Trionyx "species 2"6 Geoffroy Saint-Hilaire 1809 softshell turtle
high-shelled, big Trionyx
 Testudines - Adocidae
Adocidae indet.4 Cope 1869 turtle
a skull (ZISP PH 1/17) referred to Lindholmemys elegans by Nessov 1986 (fig. 11). Danilov & Parham (2005) "hypothesize that ZISP PH 1/17 is the skull of either Shachemys ancestralis or “Adocus” aksary"
Adocus aksary6 Nessov and Krasovskaya 1984 turtle
cf. Shachemydinae sp.6 Khosatzky 1977 turtle
 Testudines - Lindholmemydidae
Lindholmemys elegans n. sp. Riabinin 1935 turtle
 Theropoda -
 Avetheropoda - Caenagnathidae
Caenagnathasia martinsoni n. gen. n. sp.3
Caenagnathasia martinsoni n. gen. n. sp.3 Currie et al. 1994 coelurosaur
Holotype. CCMGE 401/12457, fused dentaries; referred material: . CCMGE 402/12457, a partial right dentary (Currie et al., 1994: fig. 12); ZIN PH 2354/16, incomplete fused dentaries; ZIN PH 932/16, ZIN PH 933/16, ZIN PO 4603, and ZIN PO 5234, cervical vertebrae; ZIN PH 934/16, ZIN PH 935/16, ZIN PH 936/16, and ZIN PH 937/16, dorsal vertebrae; ZIN PH 802/16 and ZIN PO 4623, partial synsacra; CCMGE 479/12457, proximal portion of femur
 Avetheropoda - Dromaeosauridae
Itemirus medullaris13 Kurzanov 1976 coelurosaur
ZIN PH 92/16
 Avetheropoda -
Tyrannosauroidea indet.2 Walker 1964 coelurosaur
ZIN PH 2296/16
? Alectrosaurus sp.9 Gilmore 1933 coelurosaur
"Segnosauria indet." = Therizinosauroidea9 Russell and Dong 1994 coelurosaur
 Ornithischia -
Ornithischia indet.9 ornithischian
 Neornithischia -
Ornithopoda indet.9 bird foot dinosaur
Levnesovia transoxiana12 Sues and Averianov 2009 bird foot dinosaur
CCMGE 566/12457
 Neornithischia - Hadrosauridae
Hadrosauridae indet. Cope 1869 hadrosaurid
or ? Ceratopsidae
 Squamata - Teiidae
Teiidae indet.9 Gray 1827 squamates
 Squamata -
? Gekkota indet.9 Cuvier 1817 squamates
Mammalia
 Tribosphenida -
Daulestes kulbeckensis n. gen. n. sp.5
Daulestes kulbeckensis n. gen. n. sp.5 Trofimov and Nessov 1979 eutherian
 Tribosphenida - Zalambdalestidae
Kulbeckia kulbecke1 Nessov 1993 eutherian
CCMGE 5/12176, M2
 Tribosphenida - Zhelestidae
"Sorlestes budan" = Zhelestes temirkazyk11, Aspanlestes aptap n. gen. n. sp.7
"Sorlestes budan" = Zhelestes temirkazyk11 Nessov 1985 eutherian
CCMGE 3/12953 (right M1)
Aspanlestes aptap n. gen. n. sp.7 Nessov 1985 eutherian
CCMGE 4/12176
 Theriiformes - Spalacotheriidae
Shalbaatar bakht n. gen. n. sp.10
Shalbaatar bakht n. gen. n. sp.10 Nessov 1997 mammal
including holotype manidble (N C.82622)